Abstract
The nomenclature of the three names that Emmanuel Drake del Castillo published in Kalanchoe (Crassulaceae subfam. Kalanchooideae), K. antanosiana, K. beharensis, and K. linearifolia, in 1903 is reviewed and discussed. Kalanchoe beharensis and K. linearifolia are today accepted species, but K. antanosiana is widely treated as a synonym of K. orgyalis. Corrections are made to the typification of the names K. beharensis and K. vantieghemii to maintain the current application of these names, with the latter being a homotypic synonym of K. beharensis.
References
Adanson, M. (1763) Familles des Plantes. Seconde Partie. Etat actuel de la Botanike. Chez Vincent, Imprimeur-Libraire de M<sup>gt</sup> le Comte de Provence, Paris, 640 pp. [https://www.biodiversitylibrary.org/item/6958#page/794/mode/1up]
Baena, S., Beentje, H., Cameron, A., Lowry, P., Moat, J., Rabehevitra, D., Rahantamalala, J., Rakotoarisoa, S., Randriarimalala, Z.L., Rasolohery, A., Schatz, G., Smith, P. & Tabor, K. (2007) Atlas of the vegetation of Madagascar. / Atlas de la vegetation de Madagascar. In: J. Moat & P. Smith (Eds.) Atlas of the vegetation of Madagascar. / Atlas de la vegetation de Madagascar. Kew Publishing, Royal Botanic Gardens, Kew, Richmond, Surrey, 124 pp.
Baillon, M.H. (1887) Liste des plantes de Madagascar (suite de la page 624). Bulletin Mensuel de la Société Linnéenne de Paris 1 (No. 84): 671–672. [https://www.biodiversitylibrary.org/page/11162078#page/24/mode/1up]
Baillon, M.H. (1888) Kalanchoe grandidieri, Plate 57 [of Plates 51–88]. In: A. Grandidier, Histoire physique, naturelle et politique Madagascar publiée par Alfred Grandidier. Vol. XXX. Histoire naturelle des plantes par M. E. Drake del Castillo. Tome II – Texte – 1, partie. 50 Fascicule. Imprimé par Autorisation de M. Le Garde des Sceaux. À l’Imprimerie Nationale, Paris, 130 plates. [https://www.biodiversitylibrary.org/item/41106#page/71/mode/1up]
Baker, J.G. (1882) Contributions to the flora of central Madagascar (continued from p. 70 and to be continued). Journal of Botany, British and Foreign. London 20: 109–114. [https://www.biodiversitylibrary.org/item/35873#page/125/mode/1up]
Boiteau, P. & Allorge-Boiteau, L. (1995) Kalanchoe (Crassulacées) de Madagascar. Systématique, écophysiologie et phytochimie. ICSN, CNRS, 91198 Gif-sur-Yvette, Éditions Karthala, Paris, 252 pp.
Berger, A. (1930) Crassulaceae. Unterfam. II. Kalanchoideae [sic Kalanchooideae]. 6. Kalanchoe. In: A. Engler & K. Prantl (Eds.) Die natürlichen Pflanzenfamilien. ed. 2, 18a. Verlag von Wilhelm Engelmann, Leipzig, pp. 402–412.
Descoings, B. (2003) Kalanchoe. In: U. Eggli (Ed.) Illustrated handbook of succulent plants: Crassulaceae. Springer Verlag, Berlin, pp. 143–181.
Dorr, L.J. (1997) Plant collectors in Madagascar and the Comoro Islands. A biographical and bibliographical guide to individuals and groups who have collected herbarium material of algae, bryophytes, fungi, lichens, and vascular plants in Madagascar and the Comoro Islands. The Trustees, Royal Botanic Gardens, Kew, 524 pp.
Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Muséum d’Histoire Naturelle 9: 35–46. [https://www.biodiversitylibrary.org/page/42893852#page/45/mode/1up]
Figueiredo, E. & Smith, G.F. (2017) (56) Request for a binding decision on the descriptive statement associated with Kalanchoe delagoensis (Crassulaceae). Taxon 66: 717. https://doi.org/10.12705/663.37
Hamet, R. (1906 [Janvier–Mars]) Note sur une nouvelle espèce de Kalanchoe [Kalanchoe vantieghemii, as ‘Van Tieghemi’]. Journal de Botanique [Morot] XX [20<sup>e</sup> [année] (N<sup>o</sup> 1–3)]: 109–111. [https://www.biodiversitylibrary.org/item/22106#page/113/mode/1up]
Hamet, R. (1907a) Monographie du genre Kalanchoe. Bulletin de l’Herbier Boissier, sér. 2, 7: 869–900. [https://www.biodiversitylibrary.org/item/105267#page/939/mode/1up]
Hamet, R. (1907b [Mars]) Note sur deux Kalanchoe malgaches. [Kalanchoe grandidieri Baill. and K. bonnieri Raym.-Hamet]. Bulletin de la Société Botanique de France [Tome Cinquante-Quatrième], Quatrième serie—Tome VII, 54: 138–139. [https://www.biodiversitylibrary.org/item/8676#page/178/mode/1up]
Hamet, R. (1908a) Monographie du genre Kalanchoe. Suite et fin. Bulletin de l’Herbier Boissier, sér. 2, 8: 17–48. [https://www.biodiversitylibrary.org/item/104945#page/31/mode/1up]
Hamet, R. (1908b) Kalanchoe luciae sp. nov. Bulletin de l’Herbier Boissier, sér. 2, 8: 254–257. [https://www.biodiversitylibrary.org/item/104945#page/290/mode/1up]
Hamet, R. (1910a [Janvier]) Sur quelques Kalanchoe peu connus. Bulletin de la Société Botanique de France [Tome Cinquante-Septième], Quatrième serie—Tome X, Quatrième series 57: 18–24. [https://www.biodiversitylibrary.org/item/8679#page/56/mode/1up]
Hamet, R. (1910b [Janvier]) Sur quelques Kalanchoe peu connus (Suite). Bulletin de la Société Botanique de France [Tome Cinquante-Septième], Quatrième serie—Tome X, Quatrième series 57: 49–54. [https://www.biodiversitylibrary.org/item/8679#page/87/mode/1up]
Hamet, R. (1910c) Kalanchoe aliciae sp. nova et K. beharensis Drake del Castillo. Bulletin de la Société Botanique de France 57: 191–194. [https://www.biodiversitylibrary.org/item/8679#page/241/mode/1up]
Hamet, R. (1912 [12 April]) Observations sur le Kalanchoe tubiflora nom. nov. Beihefte zum Botanischen Centralblatt 29 (Heft 1): 41–44. [https://www.biodiversitylibrary.org/item/27107#page/523/mode/1up]
Harvey, W.H. (1862) Order LIII. Crassulaceae, D.C. VIII. Kalanchoe, Adans. In: W.H. Harvey & O.W. Sonder (Eds.) Flora capensis [being a systematic description of the plants of the Cape Colony, Caffraria, & Port Natal]. Vol. 2. L. Reeve & Co., Ltd, Kent, pp. 378–380. [https://www.biodiversitylibrary.org/item/15233#page/386/mode/1up]
Raymond-Hamet. (1915) Sur quelques Kalanchoe de la flore Malgache. I. Bryophyllum crenatum Baker, Kalanchoe laxiflora Baker, Kalanchoe Tieghemi Raymond-Hamet et Kalanchoe crenata Raymond-Hamet. II. Kalanchoe orgyalis Baker et Kalanchoe antanosiana Drake del Castillo. III. Kitchingia porphyrocalyx Baker, Kalanchoe porphyrocalyx Baillon et Kalanchoe sulphurea Baker. IV. Kitchingia campanulata Baker, Kitchingia parviflora Baker, Kitchingia panduriformis Baker, Kitchingia amplexicaulis Baker, Kalanchoe campanulata Baillon. Kalanchoe parviflora Baillon, Kalanchoe panduriformis Baillon et Kalanchoe amplexicaulis Baillon. Annales du Musée Colonial de Marseille, sér. 3, 3: 123–164. [https://www.biodiversitylibrary.org/item/22992#page/145/mode/1up]
Raymond-Hamet & Marnier-Lapostolle, J. (1964) Le genre Kalanchoe au Jardin Botanique “Les Cèdres”. Archives du Muséum National d’Histoire Naturelle Septième Série, Tome VIII: 1–110; Plates I–XXXVII. [https://www.biodiversitylibrary.org/item/280857#page/21/mode/1up]
Scott Elliot, G.F. (1891 [22 August]) New and little known Madagascar plants, collected and enumerated by G.F. Scott Elliot, M.A., B.Sc. F.L.S. The Journal of the Linnean Society, Botany XXIX(197): 1–67, with Plates I–XII. [https://www.biodiversitylibrary.org/item/8386#page/9/mode/1up]
Smith, G.F. (2020) The kalanchoes (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1890–1972). Bradleya 38: 120–140. https://doi.org/10.25223/brad.n38.2020.a15
Smith, G.F. & Figueiredo, E. (2022) Clarifying the typification of the name Kalanchoe orgyalis (Crassulaceae subfam. Kalanchooideae). Phytotaxa 543 (1): 99–102. https://doi.org/10.11646/phytotaxa.543.1.11
Smith, G.F., Figueiredo, E. & Walker, C.C. (2021) Kalanchoe beharensis. Crassulaceae subfam. Kalanchooideae. Madagascar. Flowering Plants of Africa 67: 50–59, plate 2365.
Tulasne, L.-R. [‘Edmond’]. (1857) Fragmentum alterum (1). Floræ madagascariensis. II. Calanchoe [sic, Kalanchoe] pp. 149–150. Annales des Sciences Naturelles; Botanique sér. 4, 8: 44–163. [https://www.biodiversitylibrary.org/item/129817#page/153/mode/1up]
Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Glashütten. [Regnum Vegetabile 159], 254 pp. https://doi.org/10.12705/Code.2018
Baena, S., Beentje, H., Cameron, A., Lowry, P., Moat, J., Rabehevitra, D., Rahantamalala, J., Rakotoarisoa, S., Randriarimalala, Z.L., Rasolohery, A., Schatz, G., Smith, P. & Tabor, K. (2007) Atlas of the vegetation of Madagascar. / Atlas de la vegetation de Madagascar. In: J. Moat & P. Smith (Eds.) Atlas of the vegetation of Madagascar. / Atlas de la vegetation de Madagascar. Kew Publishing, Royal Botanic Gardens, Kew, Richmond, Surrey, 124 pp.
Baillon, M.H. (1887) Liste des plantes de Madagascar (suite de la page 624). Bulletin Mensuel de la Société Linnéenne de Paris 1 (No. 84): 671–672. [https://www.biodiversitylibrary.org/page/11162078#page/24/mode/1up]
Baillon, M.H. (1888) Kalanchoe grandidieri, Plate 57 [of Plates 51–88]. In: A. Grandidier, Histoire physique, naturelle et politique Madagascar publiée par Alfred Grandidier. Vol. XXX. Histoire naturelle des plantes par M. E. Drake del Castillo. Tome II – Texte – 1, partie. 50 Fascicule. Imprimé par Autorisation de M. Le Garde des Sceaux. À l’Imprimerie Nationale, Paris, 130 plates. [https://www.biodiversitylibrary.org/item/41106#page/71/mode/1up]
Baker, J.G. (1882) Contributions to the flora of central Madagascar (continued from p. 70 and to be continued). Journal of Botany, British and Foreign. London 20: 109–114. [https://www.biodiversitylibrary.org/item/35873#page/125/mode/1up]
Boiteau, P. & Allorge-Boiteau, L. (1995) Kalanchoe (Crassulacées) de Madagascar. Systématique, écophysiologie et phytochimie. ICSN, CNRS, 91198 Gif-sur-Yvette, Éditions Karthala, Paris, 252 pp.
Berger, A. (1930) Crassulaceae. Unterfam. II. Kalanchoideae [sic Kalanchooideae]. 6. Kalanchoe. In: A. Engler & K. Prantl (Eds.) Die natürlichen Pflanzenfamilien. ed. 2, 18a. Verlag von Wilhelm Engelmann, Leipzig, pp. 402–412.
Descoings, B. (2003) Kalanchoe. In: U. Eggli (Ed.) Illustrated handbook of succulent plants: Crassulaceae. Springer Verlag, Berlin, pp. 143–181.
Dorr, L.J. (1997) Plant collectors in Madagascar and the Comoro Islands. A biographical and bibliographical guide to individuals and groups who have collected herbarium material of algae, bryophytes, fungi, lichens, and vascular plants in Madagascar and the Comoro Islands. The Trustees, Royal Botanic Gardens, Kew, 524 pp.
Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). Bulletin du Muséum d’Histoire Naturelle 9: 35–46. [https://www.biodiversitylibrary.org/page/42893852#page/45/mode/1up]
Figueiredo, E. & Smith, G.F. (2017) (56) Request for a binding decision on the descriptive statement associated with Kalanchoe delagoensis (Crassulaceae). Taxon 66: 717. https://doi.org/10.12705/663.37
Hamet, R. (1906 [Janvier–Mars]) Note sur une nouvelle espèce de Kalanchoe [Kalanchoe vantieghemii, as ‘Van Tieghemi’]. Journal de Botanique [Morot] XX [20<sup>e</sup> [année] (N<sup>o</sup> 1–3)]: 109–111. [https://www.biodiversitylibrary.org/item/22106#page/113/mode/1up]
Hamet, R. (1907a) Monographie du genre Kalanchoe. Bulletin de l’Herbier Boissier, sér. 2, 7: 869–900. [https://www.biodiversitylibrary.org/item/105267#page/939/mode/1up]
Hamet, R. (1907b [Mars]) Note sur deux Kalanchoe malgaches. [Kalanchoe grandidieri Baill. and K. bonnieri Raym.-Hamet]. Bulletin de la Société Botanique de France [Tome Cinquante-Quatrième], Quatrième serie—Tome VII, 54: 138–139. [https://www.biodiversitylibrary.org/item/8676#page/178/mode/1up]
Hamet, R. (1908a) Monographie du genre Kalanchoe. Suite et fin. Bulletin de l’Herbier Boissier, sér. 2, 8: 17–48. [https://www.biodiversitylibrary.org/item/104945#page/31/mode/1up]
Hamet, R. (1908b) Kalanchoe luciae sp. nov. Bulletin de l’Herbier Boissier, sér. 2, 8: 254–257. [https://www.biodiversitylibrary.org/item/104945#page/290/mode/1up]
Hamet, R. (1910a [Janvier]) Sur quelques Kalanchoe peu connus. Bulletin de la Société Botanique de France [Tome Cinquante-Septième], Quatrième serie—Tome X, Quatrième series 57: 18–24. [https://www.biodiversitylibrary.org/item/8679#page/56/mode/1up]
Hamet, R. (1910b [Janvier]) Sur quelques Kalanchoe peu connus (Suite). Bulletin de la Société Botanique de France [Tome Cinquante-Septième], Quatrième serie—Tome X, Quatrième series 57: 49–54. [https://www.biodiversitylibrary.org/item/8679#page/87/mode/1up]
Hamet, R. (1910c) Kalanchoe aliciae sp. nova et K. beharensis Drake del Castillo. Bulletin de la Société Botanique de France 57: 191–194. [https://www.biodiversitylibrary.org/item/8679#page/241/mode/1up]
Hamet, R. (1912 [12 April]) Observations sur le Kalanchoe tubiflora nom. nov. Beihefte zum Botanischen Centralblatt 29 (Heft 1): 41–44. [https://www.biodiversitylibrary.org/item/27107#page/523/mode/1up]
Harvey, W.H. (1862) Order LIII. Crassulaceae, D.C. VIII. Kalanchoe, Adans. In: W.H. Harvey & O.W. Sonder (Eds.) Flora capensis [being a systematic description of the plants of the Cape Colony, Caffraria, & Port Natal]. Vol. 2. L. Reeve & Co., Ltd, Kent, pp. 378–380. [https://www.biodiversitylibrary.org/item/15233#page/386/mode/1up]
Raymond-Hamet. (1915) Sur quelques Kalanchoe de la flore Malgache. I. Bryophyllum crenatum Baker, Kalanchoe laxiflora Baker, Kalanchoe Tieghemi Raymond-Hamet et Kalanchoe crenata Raymond-Hamet. II. Kalanchoe orgyalis Baker et Kalanchoe antanosiana Drake del Castillo. III. Kitchingia porphyrocalyx Baker, Kalanchoe porphyrocalyx Baillon et Kalanchoe sulphurea Baker. IV. Kitchingia campanulata Baker, Kitchingia parviflora Baker, Kitchingia panduriformis Baker, Kitchingia amplexicaulis Baker, Kalanchoe campanulata Baillon. Kalanchoe parviflora Baillon, Kalanchoe panduriformis Baillon et Kalanchoe amplexicaulis Baillon. Annales du Musée Colonial de Marseille, sér. 3, 3: 123–164. [https://www.biodiversitylibrary.org/item/22992#page/145/mode/1up]
Raymond-Hamet & Marnier-Lapostolle, J. (1964) Le genre Kalanchoe au Jardin Botanique “Les Cèdres”. Archives du Muséum National d’Histoire Naturelle Septième Série, Tome VIII: 1–110; Plates I–XXXVII. [https://www.biodiversitylibrary.org/item/280857#page/21/mode/1up]
Scott Elliot, G.F. (1891 [22 August]) New and little known Madagascar plants, collected and enumerated by G.F. Scott Elliot, M.A., B.Sc. F.L.S. The Journal of the Linnean Society, Botany XXIX(197): 1–67, with Plates I–XII. [https://www.biodiversitylibrary.org/item/8386#page/9/mode/1up]
Smith, G.F. (2020) The kalanchoes (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1890–1972). Bradleya 38: 120–140. https://doi.org/10.25223/brad.n38.2020.a15
Smith, G.F. & Figueiredo, E. (2022) Clarifying the typification of the name Kalanchoe orgyalis (Crassulaceae subfam. Kalanchooideae). Phytotaxa 543 (1): 99–102. https://doi.org/10.11646/phytotaxa.543.1.11
Smith, G.F., Figueiredo, E. & Walker, C.C. (2021) Kalanchoe beharensis. Crassulaceae subfam. Kalanchooideae. Madagascar. Flowering Plants of Africa 67: 50–59, plate 2365.
Tulasne, L.-R. [‘Edmond’]. (1857) Fragmentum alterum (1). Floræ madagascariensis. II. Calanchoe [sic, Kalanchoe] pp. 149–150. Annales des Sciences Naturelles; Botanique sér. 4, 8: 44–163. [https://www.biodiversitylibrary.org/item/129817#page/153/mode/1up]
Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Glashütten. [Regnum Vegetabile 159], 254 pp. https://doi.org/10.12705/Code.2018