Abstract
Amanita pallidoverruca, a new species of A. sect. Validae, is described from the Hengduan Mountains, southwestern China. This species is associated with Picea trees and is characterized by a greyish brown pileus decorated with dirty white to greyish warts, non-striate and non-appendiculate pileal margin, subapical annulus with a greyish yellow to olivaceous edge, and amyloid, ellipsoid basidiospores measuring 8–10 × 6–7 μm. Phylogenetic analyses based on nrLSU, ITS, rpb2 and tef1-α indicated that it is closely related to A. excelsa. Illustrated morphological descriptions, and comparisons with closely related and similar species are provided.
References
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<p>Bresinsky, A. & Besl, H. (1990) <em>A colour atlas of poisonous fungi: a handbook for pharmacists, doctors, and biologists. </em>Wolf Publishing Ltd., London.</p>
<p>Cai, Q., Tulloss, R.E., Tang, L.P., Tolgor, B., Zhang, P., Chen, Z.H. & Yang, Z.L. (2014) Multi-locus phylogeny of lethal amanitas: implications for species diversity and historical biogeography. <em>BMC Evolutionary Biology</em> 14: 143. https://doi.org/10.1186/1471-2148-14-143</p>
<p>Castresana, J. (2000) Selection of conserved blocks from multiple alignments for their use in phylogenetic analysis. <em>Molecular Biology and Evolution</em> 17: 540–552. https://doi.org/10.1093/oxfordjournals.molbev.a026334</p>
<p>Cho, H.J., Park, M.S., Lee, H., Oh, S.Y., Jang, Y., Fong, J.J. & Lim, Y.W. (2015) Four new species of <em>Amanita</em> in Inje County, Korea. <em>Mycobiology</em> 43: 408–414. https://doi.org/10.5941/Myco.2015.43.4.408</p>
<p>Codjia, J.E.I., Cai, Q., Zhou, S.W., Luo, H., Ryberg, M., Yorou, N.S. & Yang, Z.L. (2020) Morphology, multilocus phylogeny, and toxin analysis reveal <em>Amanita albolimbata</em>, the first lethal <em>Amanita</em> species from Benin, West Africa. <em>Frontiers in Microbiology</em> 11: 599047. https://doi.org/10.3389/fmicb.2020.599047</p>
<p>Cui, Y.Y., Cai, Q., Tang, L.P., Liu, J.W. & Yang, Z.L. (2018) The family Amanitaceae: molecular phylogeny, higher-rank taxonomy and the species in China. <em>Fungal Diversity</em> 91: 5–230. https://doi.org/10.1007/s13225-018-0405-9</p>
<p>Cui, Y.Y., Cai, Q. & Yang, Z.L. (2021) <em>Amanita chuformis</em>, a new <em>Amanita</em> species with a marginate basal bulb. <em>Mycoscience</em> 62: 29–35. https://doi.org/10.47371/mycosci.2020.08.003</p>
<p>Doyle, J.J. & Doyle, J.L. (1987) A rapid DNA isolation procedure for small quantities of fresh leaf tissue. <em>Phytochemical Bulletin</em> 19: 11–15.</p>
<p>Fraiture, A., Amalfi, M., Raspe, O., Kaya, E., Akata, I. & Degreef, J. (2019) Two new species of <em>Amanita</em> sect. <em>Phalloideae</em> from Africa, one of which is devoid of amatoxins and phallotoxins. <em>Mycokeys</em> 53: 93–125. https://doi.org/10.3897/mycokeys.53.34560</p>
<p>Fries, E.M. (1821) <em>Systema mycologicum, sistens fungorum ordines, genera et species, huc usque cognitas, quas ad normam methodi naturalis determinavit, disposuit atque descripsit </em>Gryphiswaldiae, Lund. https://doi.org/10.5962/bhl.title.5378</p>
<p>Fries, E.M. (1854) <em>Monographia Amanitarum Sueciae</em>. C.A. Leffler, Upsaliae.</p>
<p>Gardes, M. & Bruns, T.D. (1993) ITS primers with enhanced specificity for basidiomycetes-application to the identification of mycorrhizae and rusts. <em>Molecular Ecology</em> 2: 113–118. https://doi.org/10.1111/j.1365-294X.1993.tb00005.x</p>
<p>Hall, T.A. (1999) BioEdit: a user-friendly biological sequence alignment editor and analysis program for Windows 95/98/NT. <em>Nucleic Acids Symposium Series</em> 41: 95–98.</p>
<p>Hongo, T. (1959) The Agaricales of Japan I-(1). <em>Memoirs of the Faculty of Liberal Arts and Education, Shiga University, Part2, Natural Science</em> 9: 47–94.</p>
<p>Imai, S. (1938) Studies in the Agaricaceae of Hokkaido. I. <em>Journal of the Faculty of Agriculture, Hokkaido Imperial University</em> 43: 1–31.</p>
<p>Imai, S. (1933) Studies on the Agaricaceae of Japan I. Volvate agarics in Hokkaido. <em>The Botanical Magazine (Tokyo)</em> 47: 423–432.</p>
<p>Jabeen, S., Kiran, M., Khan, J., Ahmad, I., Ahmad, H., Sher, H. & Khalid, A.N. (2019) <em>Amanita ahmadii</em>, a new species of <em>Amanita </em>subgenus <em>Amanitina</em> section<em> Validae</em> from Pakistan. <em>MycoKeys</em> 56: 81–99. https://doi.org/10.3897/mycokeys.56.31819</p>
<p>Jin, Y.J., Yoo, K.B., Hong, S.B., Kwon, S.W., Kim, S.J., Kim, S.H. & Seok, S.J. (2015) Notes on the unrecorded species of subgenus <em>Lepidella</em> in Korean genus <em>Amanita</em>. <em>The Korean Journal of Mycology</em> 43: 216–223.</p>
<p>Jo, J.W., Kwag, Y.N., Kim, H.S., Lee, H., Han, S.K., Han, J.G., Oh, S.H. & Kim, C.S. (2020) A new species and two new records of <em>Amanita</em> (Amanitaceae; Basidiomycota) from South Korea. <em>Phytotaxa</em> 451: 21–33. https://doi.org/10.11646/phytotaxa.451.1.2</p>
<p>Katoh, K. & Standley, D.M. (2013) MAFFT multiple sequence alignment software version 7: improvements in performance and usability. <em>Molecular Biology and Evolution</em> 30: 772–780. https://doi.org/10.1093/molbev/mst010</p>
<p>Kornerup, A. & Wanscher, J.H. (1981) <em>Taschenlexikon der Farben. 3. Aufl. </em>Muster-Schmidt Verlag, Göttingen.</p>
<p>Lanfear, R., Calcott, B., Ho, S.Y.W. & Guindon, S. (2012) PartitionFinder: combined selection of partitioning schemes and substitution models for phylogenetic analyses. <em>Molecular Biology and Evolution</em> 29: 1695–1701. https://doi.org/10.1093/molbev/mss020</p>
<p>Mehmood, T., Das, K., Hosen, M.I., Bhatt, R.P., Uniyal, P. & Singh, U. (2018) Two new species of <em>Amanita</em> (Amanitaceae) from North-western Himalaya, India. <em>Phytotaxa</em> 367: 219–232. https://doi.org/10.11646/phytotaxa.367.3.2</p>
<p>Mighell, K.S., Henkel, T.W., Koch, R.A., Chin, M.L., Brann, M.A. & Aime, M.C. (2021) <em>Amanita</em> in the Guineo-Congolian rainforest: epitypes and new species from the Dja Biosphere Reserve, Cameroon. <em>Mycologia </em>113: 168–190. https://doi.org/10.1080/00275514.2020.1816386</p>
<p>Mighell, K.S., Henkel, T.W., Koch, R.A., Goss, A. & Aime, M.C. (2019) New species of <em>Amanita</em> subgen.<em> Lepidella</em> from Guyana. <em>Fungal Systematics and Evolution</em> 3: 1–12. https://doi.org/10.3114/fuse.2019.03.01</p>
<p>Mu, M., Huang, H.Y., Zhang, W.H. & Tang, L.P. (2021) A new species with pink lamellae of <em>Amanita </em>section <em>Caesareae</em> from China. <em>Phytotaxa</em> 478: 141–150. https://doi.org/10.11646/phytotaxa.478.1.10</p>
<p>Nascimento, C.C., Sa, M.C.A., Bezerra, J.L. & Wartchow, F. (2018) <em>Amanita dulciodora</em> (Amanitaceae, Basidiomycota), a striking new species of <em>Amanita</em> section <em>Lepidella </em>from Northeast Brazil. <em>Plant Ecology and Evolution</em> 151: 262–270. https://doi.org/10.5091/plecevo.2018.1395</p>
<p>Neville, P. & Poumarat, S. (2004) <em>Amaniteae: Amanita, Limacella & Torrendia. </em>Edizioni Candusso, Alssio.</p>
<p>Persoon, C.H. (1797) <em>Tentamen dispositionis methodicae fungorum in classes, ordines, genera et familias. </em>Lipsiae, P.P. Wolf. https://doi.org/10.5962/bhl.title.42674</p>
<p>Quélet, L. (1872) Les Champignons du Jura et des Vosges. <em>Mémoires de la Société d’Émulation de Montbéliard </em>sér. II 5: 43–332.</p>
<p>Rehner, S.A. & Buckley, E. (2005) A <em>Beauveria</em> phylogeny inferred from nuclear ITS and EF1-alpha sequences: evidence for cryptic diversification and links to<em> Cordyceps teleomorphs</em>. <em>Mycologia</em> 97: 84–98. https://doi.org/10.3852/mycologia.97.1.84</p>
<p>Ronquist, F. & Huelsenbeck, J.P. (2003) MrBayes 3: Bayesian phylogenetic inference under mixed models. <em>Bioinformatics</em> 19: 1572–1574. https://doi.org/10.1093/bioinformatics/btg180</p>
<p>Saba, M., Haelewaters, D., Fiaz, M., Khalid, A.N. & Pfister, D.H. (2019) <em>Amanita mansehraensis</em>, a new species in section <em>Vaginatae</em> from Pakistan. <em>Phytotaxa</em> 409: 189–201. https://doi.org/10.11646/phytotaxa.409.4.1</p>
<p>Saccardo, P.A. (1887) Sylloge hymenomycetum, vol. I. Agaricineae. <em>Sylloge fungorum omnium hucusque cognitorum</em> 5: 1–1146.</p>
<p>Semwal, K.C., Das, K., Bhatt, R.P., Mehmood, T. & Bhatt, V.K. (2020) <em>Amanita pseudorufobrunnescens</em>, a new species of <em>Amanita</em> section <em>Amidella</em> from Indian Himalaya. <em>Kew Bulletin</em> 75: 22. https://doi.org/10.1007/s12225-020-9874-6</p>
<p>Smith, S.A. & Dunn, C.W. (2008) Phyutility: a phyloinformatics tool for trees, alignments and molecular data. <em>Bioinformatics</em> 24: 715–716. https://doi.org/10.1093/bioinformatics/btm619</p>
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