A revised phylogenetic classification of tribe Phyllantheae (Phyllanthaceae)

RODERICK W. BOUMAN; PAUL J.A. KEBLER; IAN R.H. TELFORD; JEREMY J. BRUHL; JOERI S. STRIJK; RICHARD M.K. SAUNDERS; HANS-JOACHIM ESSER; BANESSA FALCÓN-HIDALGO; PETER C. VAN WELZEN

doi:10.11646/phytotaxa.540.1.1

Issue: Vol. 540 No. 1: 18 March 2022

Type: Monograph

Published: 2022-03-18

DOI: 10.11646/phytotaxa.540.1.1

Page range: 1-100

Abstract views: 548

PDF downloaded: 22

A revised phylogenetic classification of tribe Phyllantheae (Phyllanthaceae)

RODERICK W. BOUMAN⁺⁻
PAUL J.A. KEBLER⁺⁻
IAN R.H. TELFORD⁺⁻
JEREMY J. BRUHL⁺⁻
JOERI S. STRIJK⁺⁻
RICHARD M.K. SAUNDERS⁺⁻
HANS-JOACHIM ESSER⁺⁻
BANESSA FALCÓN-HIDALGO⁺⁻
PETER C. VAN WELZEN⁺⁻

Naturalis Biodiversity Center, PO Box 9517, 2300 RA Leiden, The Netherlands; Hortus botanicus Leiden, Leiden University, PO Box 9500, 2300 RA Leiden, The Netherlands; Institute of Biology Leiden, Leiden University, PO Box 9505, 2300 RA Leiden, The Netherlands

Hortus botanicus Leiden, Leiden University, PO Box 9500, 2300 RA Leiden, The Netherlands; Institute of Biology Leiden, Leiden University, PO Box 9505, 2300 RA Leiden, The Netherlands

Botany and N. C. W. Beadle Herbarium, School of Environmental and Rural Science, University of New England, Armidale, NSW 2351, Australia

Institute for Biodiversity and Environmental Research, Jalan Tungku Link, Universiti Brunei Darussalam, Brunei Darussalam; Alliance for Conservation Tree Genomics, Pha Tad Ke Botanical Garden, PO Box 959, 06000 Luang Prabang, Lao PDR

Division of Ecology & Biodiversity, School of Biological Sciences, The University of Hong Kong, Pokfulam Road, Hong Kong, China

Botanische Staatssammlung München, Menzinger Strasse 67, D-80638 München (Munich), Germany

Jardin Botánico Nacional, Unisversidad de La Habana, Carretera “El Rocío” km 3.5, Calabazar, Boyeros, La Habana, Cuba

Naturalis Biodiversity Center, PO Box 9517, 2300 RA Leiden, The Netherlands; Institute of Biology Leiden, Leiden University, PO Box 9505, 2300 RA Leiden, The Netherlands

Phyllanthus paraphyletic Malpighiales Phyllanthaceae Phyllantheae Eudicots

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Abstract

The majority of tribe Phyllantheae (Phyllanthaceae) is currently placed in the paraphyletic genus Phyllanthus and discussions have persisted on how to resolve this issue. Here, we split Phyllanthus into ten monophyletic genera, which are all reinstatements of former genera, but with changes made to the circumscription and constituent species of each group. The genera Breynia, Glochidion and Synostemon were recently found to be nested within Phyllanthus and discussions ensued whether or not to subsume everything into Phyllanthus s.l. Instead of combining all these genera, we here implement the solution of splitting Phyllanthus into strictly monophyletic genera to ensure that the classification is consistent with the latest phylogenetic results. The new classification is based on a phylogenetic framework combined with differences in habit, branching type, floral, fruit and pollen morphology. With this new division of the genus Phyllanthus, tribe Phyllantheae will consist of the following 18 genera: Breynia, Cathetus, Cicca, Dendrophyllanthus, Emblica, Flueggea, Glochidion, Heterosavia, Kirganelia, Lingelsheimia, Lysiandra, Margaritaria, Moeroris, Nellica, Nymphanthus, Phyllanthus, Plagiocladus and Synostemon. As a result of the reinstated genera, five new names for illegitimate combinations or previous overlooked nomenclatural anomalies and 645 new combinations are proposed. Several keys are provided to distinguish the reinstated genera. Full species lists are given for the reinstated genera treated here except for Breynia, Synostemon and Glochidion.

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