Abstract
Citronella suaveolens is reported from Quang Nam Province in southern Vietnam, representing the first record of this species and the genus Citronella in the country and in mainland Southeast Asia in general. Detailed photographic illustrations of studied specimens are provided. A key to the Vietnamese genera and species of Cardiopteridaceae is presented. Diversity of Citronella in the Malesian region is discussed. Floristic affinities of Malesia and Eastern Indochina are highlighted by a list of illustrative examples.
References
APG IV (2016) An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV. Botanical Journal of the Linnean Society 181 (1): 1–20. https://doi.org/10.1111/boj.12385
Averyanov, L.V. (2010) Species of orchids (Orchidaceae) newly recorded to the flora of Vietnam. Taiwania 55 (1): 1–7. https://doi.org/10.6165/tai.2010.55(1).1
Averyanov, L.V. (2012a) New orchid taxa and records in the flora of Vietnam. Taiwania 57 (2): 127–152. https://doi.org/10.6165/10.6165/tai.2012.57(2).127
Averyanov, L.V. (2012b) New orchids (Orchidaceae) in the flora of Vietnam. Turczaninowia 15 (1): 11–18.
Averyanov, L.V., Duy, N.V. & Phan, K.L. (2012) Hymenorchis phitamii (Orchidaceae) – new genus and species in the flora of Vietnam. Taiwania 57 (4): 372–376. https://doi.org/10.6165/tai.2012.57.372
Averyanov, L.V., Nguyen, K.S., Nguyen, T.T., Nguyen, P.T., Nong, V.D., Nguyen, V.C. & Xuan, C.C. (2015) New orchids in the flora of Vietnam. Wulfenia 22: 137–188.
Averyanov, L.V., Nong, V.D., Nguyen, K.S., Maisak, T.V., Nguyen, V.C., Phan, Q.T., Nguyen, P.T., Nguyen, T.T. & Truong, B.V. (2016) New species of orchids (Orchidaceae) in the flora of Vietnam. Taiwania 61 (4): 319–354. https://doi.org/10.6165/tai.2016.61.319
Averyanov, L.V., Nguyen, V.C., Truong, B.V., Maisak, T.V., Luu, H.T., Nguyen, K.S., Dinh, Q.D., Nguyen, H.T., Chu, X.C., Tran, G., Nguyen, V.K. & Le, H.S. (2018a) New orchids (Orchidaceae: Cymbidieae and Vandeae) in the flora of Vietnam. Taiwania 63 (2): 119–138. https://doi.org/10.6165/tai.2018.63.119
Averyanov, L.V., Nguyen, V.C., Nguyen, H.T., Truong, B.V., Nguyen, P.T., Nguyen, K.S., Maisak, T.V., Nguyen, H.T., Bui, D.N. & Chu, X.C. (2018b) New orchids (Orchidaceae: Epidendroideae and Vandoideae) in the flora of Vietnam. Taiwania 63 (3): 195–219. https://doi.org/10.6165/tai.2018.63.195
Averyanov, L.V., Nong, V.D., Nguyen, H.T., Nuraliev, M.S., Maisak, T.V. & Nguyen, C.A. (2018c) New species of Bulbophyllum (Orchidaceae) in the flora of Vietnam. Phytotaxa 369 (1): 1–14. https://doi.org/10.11646/phytotaxa.369.1.1
Averyanov, L.V., Nguyen, V.D., Nguyen, K.S., Dinh, Q.D. & Maisak, T.V. (2018d) Silvorchis vietnamica (Orchidaceae, Orchidoideae, Vietorchidinae), a new miniature mycotrophic species from Southern Vietnam. Nordic Journal of Botany 36 (7): e01883. https://doi.org/10.1111/njb.01883
Averyanov, L.V., Nguyen, V.C., Nguyen, K.S., Maisak, T.V. & Truong, B.V. (2019) New orchids (Orchidaceae) in the flora of Vietnam I. Epidendroideae. Taiwania 64 (2): 176–188. https://doi.org/10.6165/tai.2019.64.176
Averyanov, L.V., ?skov, ?.K., Truong, B.V., Nguyen V.C., Prilepsky, N.G., Nuraliev, M.S., Maisak, T.V. & Manh, V. (2020) The genus Octarrhena (Orchidaceae, Thelasiinae) in Vietnam with description of two new species. Phytotaxa 459 (4): 265–275. https://doi.org/10.11646/phytotaxa.459.4.2
Backer, C.A. & Bakhuizen van den Brink, R.C. (1965) Flora of Java. Vol. 2. N.V.P. Noordhoff, Groningen, 641 pp.
Beaman, J.H., Anderson, C. & Beaman, R.S. (2001) The plants of Mount Kinabalu. 4. Dicotyledon families Acanthaceae to Lythraceae. Natural History Publications (Borneo), Kota Kinabalu, 750 pp.
Blume, C.L. (1850) Ord. Olacineae. In: Blume, C.L. (Ed.) Museum botanicum Lugduno-Batavum, sive, Stirpium exoticarum novarum vel minus cognitarum ex vivis aut siccis brevis expositio et description 1 (16). E.J. Brill, Lugduno-Batavia, pp. 245–251. https://doi.org/10.5962/bhl.title.274
Chen, C.W., Hsu, T.C., Chao, Y.S., Lu, P.F., Li, C.W., Tram, N.K.T., Nguyen, I.L.P.T., Dang, M.T., Luu, H.T. & Parris, B. (2020) A newly recorded genus and twelve newly recorded species of ferns for Vietnam from Lang Biang Plateau. Phytotaxa 443: 121–143. https://doi.org/10.11646/phytotaxa.443.2.1
Christenhusz, M.J., Fay, M.F. & Chase, M.W. (2017) Plants of the world: an illustrated encyclopedia of vascular plants. The University of Chicago Press, Chicago, 792 pp. https://dx.doi.org/10.7208/chicago/9780226536705.001.0001
Dang, M.Q., Ormerod, P., Bui, V.H., Truong, Q.T., Duong, M.T.T.N., Dang, V.S. & Truong, B.V. (2021) Mycaranthes latifolia (Orchidaceae), a new record for Vietnam. Rheedea 31 (3): 206–210. https://dx.doi.org/10.22244/rheedea.2021.31.03.12
Don, D. (1832) On the character and affinities of certain genera, chiefly belonging to the Flora Peruviana. The Edinburgh New Philosophical Journal 13: 233–244.
Gagnepain, F. (1910) Malvacée, Tiliacée, Santalacée et Olacacées nouvelles. Notulae Systematicae 1 (7): 194–206.
Gagnepain, F. (1911) Cardioptéridacées. In: Lecomte, H. (Ed.) Flore Générale de L’Indochine. Vol. 1, fasc. 7. Masson et C<sup>ie</sup>, Paris, pp. 846–849. https://doi.org/10.5962/bhl.title.44886
Hasskarl, J.C. (1843) Adnotationes de plantis quibusdam Javanicis nonnullisque Japonicis, haud rite cognitis, e Catalogo Horti Bogoriensis. Accedunt nonnullae novae species. Tijdschrift voor Natuurlijke Geschiedenis en Physiologie 10: 115–150.
Hasskarl, J.C. (1855) Retzia sive observationes botanicae, quas in primis in Horto Botanico Bogoriensi mensibus Februario ad Juliaum 1855 fecit J.K. Hasskarl. Pugillus Primus. Lange & Co., Batavia, 252 pp.
Howard, R.A. (1940) Studies of the Icacinaceae. I. Preliminary taxonomic notes. Journal of the Arnold Arboretum 21 (4): 461–489.
Howard, R.A. (1942) Studies of the Icacinaceae. V. A revision of the genus Citronella D. Don. Contributions from the Gray Herbarium of Harvard University 142: 60–89.
Hsu, T.C., Chen, C.W., Hung, H.C., Tram, N.K.T., Truong, Q.C., Luu, H.T. & Li, C.W. (2020) New and noteworthy orchids (Orchidaceae) discovered in Langbiang Plateau, southern Vietnam 1. Taiwania 65 (2): 237–248. https://doi.org/10.6165/tai.2020.65.237
Kårehed, J. (2001) Multiple origin of the tropical forest tree family Icacinaceae. American Journal of Botany 88 (12): 2259–2274. https://doi.org/10.2307/3558388
Kurz, S. (1870) On some new or imperfectly known Indian plants. Journal of the Asiatic Society of Bengal 39 (2): 61–91.
Li, H.T., Luo, Y., Gan, L., Ma, P.F., Gao, L.M., Yang, J.B., Cai, J., Gitzendanner, M.A., Fritsch, P.W., Zhang, T., Jin, J.J., Zeng, C.X., Wang, H., Yu, W.B., Zhang, R., van der Bank, M., Olmstead, R.G., Hollingsworth, P.M., Chase, M.W. & Soltis, D.E. (2021) Plastid phylogenomic insights into relationships of all flowering plant families. BMC Biology 19: 232. https://doi.org/10.1186/s12915-021-01166-2
Long, D.G. (1991) Family 107. Cardiopteridaceae. In: Grierson, A.J.C. & Long, D.G. (Eds.) Flora of Bhutan, including a record of plants from Sikkim, volume 2, part 1. Royal Botanic Garden, Edinburgh, pp. 106.
Lu, N.T., Viet, H.N., Sennikov, A.N., Averyanov, L.V., Do, T.V., Kuznetsov, A.N., Kuznetsova, S.P. & Nuraliev, M.S. (2020) Checklist of Lycopodiaceae in Vietnam with three new records and one lectotypification. Phytotaxa 452 (1): 19–32. https://doi.org/10.11646/phytotaxa.452.1.2
Merrill, E.D. (1919) New or noteworthy Philippine plants, XV. The Philippine Journal of Science 14 (4): 365–457.
Miers, J. (1852) On some genera of the Icacinaceae. The Annals and Magazine of Natural History, series 2, 10: 108–119.
Miquel, F.A.W. (1861) Flora van Nederlandsch Indie. Eerste Bijvoegsel. Van der Post, Amsterdam, 656 pp. https://doi.org/10.5962/bhl.title.93
Munzinger, J. & Levionnois, S. (2016) Novitates neocaledonicae III: A new species of Citronella (Cardiopteridaceae) endemic to New Caledonia. Phytotaxa 245 (3): 223–228. http://dx.doi.org/10.11646/phytotaxa.245.3.5
Naiki, A., Tagane, S., Chhang, P., Toyama, H., Zhu, H., Dang, V.S. & Yahara, T. (2015) Flora of Bokor National Park, Cambodia II: four new species and nine new records of Lasianthus (Rubiaceae) from Cambodia. Acta Phytotaxonomica et Geobotanica 66 (3): 153–179. https://doi.org/10.18942/apg.KJ00010115702
Nguyen Tien Ban (2003) 164. Icacinaceae Miers, 1851. In: Nguyen Tien Ban (Ed.) Checklist of plant species of Vietnam. Vol. 2. Agriculture Publishing House, Hanoi, pp. 1109–1115. [in Vietnamese]
Nuraliev, M.S., Averyanov, L.V., Kuznetsov, A.N. & Kuznetsova, S.P. (2015) Review of the genus Plocoglottis (Orchidaceae) in Cambodia, Laos and Vietnam. Wulfenia 22: 189?199.
Nuraliev, M.S., Zhang, D., Kuznetsov, A.N. & Kuznetsova, S.P. (2018) Two new records of non-photosynthetic Burmannia species (Burmanniaceae) from Laos and Vietnam. Wulfenia 25: 52–56.
Nuraliev, M.S., Remizowa, M.V. & Sokoloff, D.D. (2020) Flower structure and development in Vietnamese Sciaphila (Triuridaceae: Pandanales): refined knowledge of the morphological misfit family and implications for taxonomy. PeerJ 8: e10205. https://doi.org/10.7717/peerj.10205
Parris, B., Chen, C.W., Hsu, T.C., Lu, N.T., Nguyen, Q.D. & Luu, H.T. (2015) New species and records of grammitid ferns (Polypodiaceae) for Vietnam. Phytotaxa 226: 39–50. http://dx.doi.org/10.11646/phytotaxa.226.1.4
Pham Hoang Ho (2003) An illustrated flora of Vietnam. Vol. 2. Youth Publishing House, Ho Chi Minh City, 951 pp. [in Vietnamese]
Remizowa, M.V., Nuraliev, M.S., Averyanov, L.V., Kuznetsov, A.N. & Kuznetsova, S.P. (2017) A revision of the family Petrosaviaceae in Vietnam. Nordic Journal of Botany 35 (3): 262–271. https://doi.org/10.1111/njb.01480
Royle, J.F. (1834) Illustrations of the botany and other branches of the natural history of the Himalayan Mountains, and of the flora of Cashmere. Wm. H. Allen & Co., London, 472 pp.
Schori, M. (2016) Cardiopteridaceae. In: Kadereit, J.W. & Bittrich, V. (Eds.) Flowering plants. Eudicots. The families and genera of vascular plants. Vol. 14. Springer, Cham, pp. 131–136. https://doi.org/10.1007/978-3-319-28534-4_8
Sleumer, H. (1969) Materials towards the knowledge of the Icacinaceae of Asia, Malesia, and adjacent areas. Blumea 17 (1): 181–264.
Sleumer, H. (1971) Icacinaceae. In: van Steenis, C.G.G.J. (Ed.) Flora Malesiana, series I, vol. 7 (1). Noordhoff, Leiden, pp. 1–87.
Stevens, P.F. (2001 onwards) Angiosperm Phylogeny Website. Version 14. Available from: http://www.mobot.org/MOBOT/research/APweb/ (accessed 22 October 2021)
Tagane, S., Wijedasa, L.S., Chhang, P., Toyama, H. & Yahara, T. (2015) Two new records for Cambodia’s forest flora, Memecylon corticosum var. kratense and M. paniculatum (Melastomataceae). Cambodian Journal of Natural History 2015 (2): 139–143.
Tagane, S., Toyama, H., Chhang, P., Dang, V.S. & Yahara, T. (2016a) New records of Xanthophyllum ellipticum and X. obscurum (Polygalaceae) in Indochina, with an identification key for species in the region. Cambodian Journal of Natural History 2016: 15–19.
Tagane, S., Naiki, A., Dang, V.S., Tran, H. & Yahara, T. (2016b) A new record of Gaertnera junghuhniana Miq. (Rubiaceae) from southern Vietnam. Acta Phytotaxonomica et Geobotanica 67 (2): 127–130. https://doi.org/10.18942/apg.201602
Tagane, S., Souladeth, P., Rueangruea, S., Okabe, N., Zhang, M., Chayer, S., Yang, C.J. & Yahara, T. (2018a) Flora of Nam Kading National Protected Area II: 30 new records of angiosperms to Laos. Edinburgh Journal of Botany 75 (1): 107–116. https://doi.org/10.1017/S0960428617000361
Tagane, S., Souladeth, P., Zhang, M. & Yahara, T. (2018b) Flora of Nam Kading National Protected Area IV: Two new species of Annonaceae, Monoon namkadingense and Neo-uvaria laosensis. Phytotaxa 336 (1): 82–88. http://dx.doi.org/10.11646/phytotaxa.336.1.6
Takhtajan, A.L. (2009) Flowering Plants. 2nd ed. Springer, New York, 871 pp. https://doi.org/10.1007/978-1-4020-9609-9
Truong, V.B., Truong, T.Q., Dang, V.S., Nguyen, T.M. & Vermeulen, J.J. (2020a) New records of Bulbophyllum in the Flora of Vietnam. Lankesteriana 20 (1): 31–36. http://dx.doi.org/10.15517/lank.v20i1.41130
Truong, V.B., Nguyen, V.C., Ormerod, P., Truong, Q.C., Chu, V.T. & Kumar, P. (2020b) The genus Corybas in Vietnam, with two new records. Lankesteriana 20 (1): 49–55. http://dx.doi.org/10.15517/lank.v20i1.41330
Utteridge, T.M.A. & Schori, M. (2011) Updating Malesian Icacinaceae. Gardens’ Bulletin Singapore 63 (1–2): 105–118.
Vislobokov, N.A., Nuraliev, M.S., Kuznetsov, A.N. & Kuznetsova, S.P. (2019) Aspidistra micrantha (Asparagaceae, Nolinoideae), a new species from Vietnam with unusually small flowers. Phytotaxa 422 (3): 289–294. https://doi.org/10.11646/phytotaxa.422.3.8
Wu, S.G., Xiang, J.Y., Phan, K.L. & Souliya, O. (2012) Some new records of ferns from Vietnam and Laos (3) Pteris (Pteridaceae). Plant Diversity & Resources 34 (1): 22–24. http://dx.doi.org/10.3724/sp.j.1143.2012.11100
Zhang, C., Zhang, T., Luebert, F., Xiang, Y., Huang, C.H., Hu, Y., Rees, M., Frohlich, M.W., Qi, J., Weigend, M. & Ma, H. (2020) Asterid phylogenomics/phylotranscriptomics uncover morphological evolutionary histories and support phylogenetic placement for numerous whole-genome duplications. Molecular Biology and Evolution 37 (11): 3188–3210. http://dx.doi.org/10.1093/molbev/msaa160
Averyanov, L.V. (2010) Species of orchids (Orchidaceae) newly recorded to the flora of Vietnam. Taiwania 55 (1): 1–7. https://doi.org/10.6165/tai.2010.55(1).1
Averyanov, L.V. (2012a) New orchid taxa and records in the flora of Vietnam. Taiwania 57 (2): 127–152. https://doi.org/10.6165/10.6165/tai.2012.57(2).127
Averyanov, L.V. (2012b) New orchids (Orchidaceae) in the flora of Vietnam. Turczaninowia 15 (1): 11–18.
Averyanov, L.V., Duy, N.V. & Phan, K.L. (2012) Hymenorchis phitamii (Orchidaceae) – new genus and species in the flora of Vietnam. Taiwania 57 (4): 372–376. https://doi.org/10.6165/tai.2012.57.372
Averyanov, L.V., Nguyen, K.S., Nguyen, T.T., Nguyen, P.T., Nong, V.D., Nguyen, V.C. & Xuan, C.C. (2015) New orchids in the flora of Vietnam. Wulfenia 22: 137–188.
Averyanov, L.V., Nong, V.D., Nguyen, K.S., Maisak, T.V., Nguyen, V.C., Phan, Q.T., Nguyen, P.T., Nguyen, T.T. & Truong, B.V. (2016) New species of orchids (Orchidaceae) in the flora of Vietnam. Taiwania 61 (4): 319–354. https://doi.org/10.6165/tai.2016.61.319
Averyanov, L.V., Nguyen, V.C., Truong, B.V., Maisak, T.V., Luu, H.T., Nguyen, K.S., Dinh, Q.D., Nguyen, H.T., Chu, X.C., Tran, G., Nguyen, V.K. & Le, H.S. (2018a) New orchids (Orchidaceae: Cymbidieae and Vandeae) in the flora of Vietnam. Taiwania 63 (2): 119–138. https://doi.org/10.6165/tai.2018.63.119
Averyanov, L.V., Nguyen, V.C., Nguyen, H.T., Truong, B.V., Nguyen, P.T., Nguyen, K.S., Maisak, T.V., Nguyen, H.T., Bui, D.N. & Chu, X.C. (2018b) New orchids (Orchidaceae: Epidendroideae and Vandoideae) in the flora of Vietnam. Taiwania 63 (3): 195–219. https://doi.org/10.6165/tai.2018.63.195
Averyanov, L.V., Nong, V.D., Nguyen, H.T., Nuraliev, M.S., Maisak, T.V. & Nguyen, C.A. (2018c) New species of Bulbophyllum (Orchidaceae) in the flora of Vietnam. Phytotaxa 369 (1): 1–14. https://doi.org/10.11646/phytotaxa.369.1.1
Averyanov, L.V., Nguyen, V.D., Nguyen, K.S., Dinh, Q.D. & Maisak, T.V. (2018d) Silvorchis vietnamica (Orchidaceae, Orchidoideae, Vietorchidinae), a new miniature mycotrophic species from Southern Vietnam. Nordic Journal of Botany 36 (7): e01883. https://doi.org/10.1111/njb.01883
Averyanov, L.V., Nguyen, V.C., Nguyen, K.S., Maisak, T.V. & Truong, B.V. (2019) New orchids (Orchidaceae) in the flora of Vietnam I. Epidendroideae. Taiwania 64 (2): 176–188. https://doi.org/10.6165/tai.2019.64.176
Averyanov, L.V., ?skov, ?.K., Truong, B.V., Nguyen V.C., Prilepsky, N.G., Nuraliev, M.S., Maisak, T.V. & Manh, V. (2020) The genus Octarrhena (Orchidaceae, Thelasiinae) in Vietnam with description of two new species. Phytotaxa 459 (4): 265–275. https://doi.org/10.11646/phytotaxa.459.4.2
Backer, C.A. & Bakhuizen van den Brink, R.C. (1965) Flora of Java. Vol. 2. N.V.P. Noordhoff, Groningen, 641 pp.
Beaman, J.H., Anderson, C. & Beaman, R.S. (2001) The plants of Mount Kinabalu. 4. Dicotyledon families Acanthaceae to Lythraceae. Natural History Publications (Borneo), Kota Kinabalu, 750 pp.
Blume, C.L. (1850) Ord. Olacineae. In: Blume, C.L. (Ed.) Museum botanicum Lugduno-Batavum, sive, Stirpium exoticarum novarum vel minus cognitarum ex vivis aut siccis brevis expositio et description 1 (16). E.J. Brill, Lugduno-Batavia, pp. 245–251. https://doi.org/10.5962/bhl.title.274
Chen, C.W., Hsu, T.C., Chao, Y.S., Lu, P.F., Li, C.W., Tram, N.K.T., Nguyen, I.L.P.T., Dang, M.T., Luu, H.T. & Parris, B. (2020) A newly recorded genus and twelve newly recorded species of ferns for Vietnam from Lang Biang Plateau. Phytotaxa 443: 121–143. https://doi.org/10.11646/phytotaxa.443.2.1
Christenhusz, M.J., Fay, M.F. & Chase, M.W. (2017) Plants of the world: an illustrated encyclopedia of vascular plants. The University of Chicago Press, Chicago, 792 pp. https://dx.doi.org/10.7208/chicago/9780226536705.001.0001
Dang, M.Q., Ormerod, P., Bui, V.H., Truong, Q.T., Duong, M.T.T.N., Dang, V.S. & Truong, B.V. (2021) Mycaranthes latifolia (Orchidaceae), a new record for Vietnam. Rheedea 31 (3): 206–210. https://dx.doi.org/10.22244/rheedea.2021.31.03.12
Don, D. (1832) On the character and affinities of certain genera, chiefly belonging to the Flora Peruviana. The Edinburgh New Philosophical Journal 13: 233–244.
Gagnepain, F. (1910) Malvacée, Tiliacée, Santalacée et Olacacées nouvelles. Notulae Systematicae 1 (7): 194–206.
Gagnepain, F. (1911) Cardioptéridacées. In: Lecomte, H. (Ed.) Flore Générale de L’Indochine. Vol. 1, fasc. 7. Masson et C<sup>ie</sup>, Paris, pp. 846–849. https://doi.org/10.5962/bhl.title.44886
Hasskarl, J.C. (1843) Adnotationes de plantis quibusdam Javanicis nonnullisque Japonicis, haud rite cognitis, e Catalogo Horti Bogoriensis. Accedunt nonnullae novae species. Tijdschrift voor Natuurlijke Geschiedenis en Physiologie 10: 115–150.
Hasskarl, J.C. (1855) Retzia sive observationes botanicae, quas in primis in Horto Botanico Bogoriensi mensibus Februario ad Juliaum 1855 fecit J.K. Hasskarl. Pugillus Primus. Lange & Co., Batavia, 252 pp.
Howard, R.A. (1940) Studies of the Icacinaceae. I. Preliminary taxonomic notes. Journal of the Arnold Arboretum 21 (4): 461–489.
Howard, R.A. (1942) Studies of the Icacinaceae. V. A revision of the genus Citronella D. Don. Contributions from the Gray Herbarium of Harvard University 142: 60–89.
Hsu, T.C., Chen, C.W., Hung, H.C., Tram, N.K.T., Truong, Q.C., Luu, H.T. & Li, C.W. (2020) New and noteworthy orchids (Orchidaceae) discovered in Langbiang Plateau, southern Vietnam 1. Taiwania 65 (2): 237–248. https://doi.org/10.6165/tai.2020.65.237
Kårehed, J. (2001) Multiple origin of the tropical forest tree family Icacinaceae. American Journal of Botany 88 (12): 2259–2274. https://doi.org/10.2307/3558388
Kurz, S. (1870) On some new or imperfectly known Indian plants. Journal of the Asiatic Society of Bengal 39 (2): 61–91.
Li, H.T., Luo, Y., Gan, L., Ma, P.F., Gao, L.M., Yang, J.B., Cai, J., Gitzendanner, M.A., Fritsch, P.W., Zhang, T., Jin, J.J., Zeng, C.X., Wang, H., Yu, W.B., Zhang, R., van der Bank, M., Olmstead, R.G., Hollingsworth, P.M., Chase, M.W. & Soltis, D.E. (2021) Plastid phylogenomic insights into relationships of all flowering plant families. BMC Biology 19: 232. https://doi.org/10.1186/s12915-021-01166-2
Long, D.G. (1991) Family 107. Cardiopteridaceae. In: Grierson, A.J.C. & Long, D.G. (Eds.) Flora of Bhutan, including a record of plants from Sikkim, volume 2, part 1. Royal Botanic Garden, Edinburgh, pp. 106.
Lu, N.T., Viet, H.N., Sennikov, A.N., Averyanov, L.V., Do, T.V., Kuznetsov, A.N., Kuznetsova, S.P. & Nuraliev, M.S. (2020) Checklist of Lycopodiaceae in Vietnam with three new records and one lectotypification. Phytotaxa 452 (1): 19–32. https://doi.org/10.11646/phytotaxa.452.1.2
Merrill, E.D. (1919) New or noteworthy Philippine plants, XV. The Philippine Journal of Science 14 (4): 365–457.
Miers, J. (1852) On some genera of the Icacinaceae. The Annals and Magazine of Natural History, series 2, 10: 108–119.
Miquel, F.A.W. (1861) Flora van Nederlandsch Indie. Eerste Bijvoegsel. Van der Post, Amsterdam, 656 pp. https://doi.org/10.5962/bhl.title.93
Munzinger, J. & Levionnois, S. (2016) Novitates neocaledonicae III: A new species of Citronella (Cardiopteridaceae) endemic to New Caledonia. Phytotaxa 245 (3): 223–228. http://dx.doi.org/10.11646/phytotaxa.245.3.5
Naiki, A., Tagane, S., Chhang, P., Toyama, H., Zhu, H., Dang, V.S. & Yahara, T. (2015) Flora of Bokor National Park, Cambodia II: four new species and nine new records of Lasianthus (Rubiaceae) from Cambodia. Acta Phytotaxonomica et Geobotanica 66 (3): 153–179. https://doi.org/10.18942/apg.KJ00010115702
Nguyen Tien Ban (2003) 164. Icacinaceae Miers, 1851. In: Nguyen Tien Ban (Ed.) Checklist of plant species of Vietnam. Vol. 2. Agriculture Publishing House, Hanoi, pp. 1109–1115. [in Vietnamese]
Nuraliev, M.S., Averyanov, L.V., Kuznetsov, A.N. & Kuznetsova, S.P. (2015) Review of the genus Plocoglottis (Orchidaceae) in Cambodia, Laos and Vietnam. Wulfenia 22: 189?199.
Nuraliev, M.S., Zhang, D., Kuznetsov, A.N. & Kuznetsova, S.P. (2018) Two new records of non-photosynthetic Burmannia species (Burmanniaceae) from Laos and Vietnam. Wulfenia 25: 52–56.
Nuraliev, M.S., Remizowa, M.V. & Sokoloff, D.D. (2020) Flower structure and development in Vietnamese Sciaphila (Triuridaceae: Pandanales): refined knowledge of the morphological misfit family and implications for taxonomy. PeerJ 8: e10205. https://doi.org/10.7717/peerj.10205
Parris, B., Chen, C.W., Hsu, T.C., Lu, N.T., Nguyen, Q.D. & Luu, H.T. (2015) New species and records of grammitid ferns (Polypodiaceae) for Vietnam. Phytotaxa 226: 39–50. http://dx.doi.org/10.11646/phytotaxa.226.1.4
Pham Hoang Ho (2003) An illustrated flora of Vietnam. Vol. 2. Youth Publishing House, Ho Chi Minh City, 951 pp. [in Vietnamese]
Remizowa, M.V., Nuraliev, M.S., Averyanov, L.V., Kuznetsov, A.N. & Kuznetsova, S.P. (2017) A revision of the family Petrosaviaceae in Vietnam. Nordic Journal of Botany 35 (3): 262–271. https://doi.org/10.1111/njb.01480
Royle, J.F. (1834) Illustrations of the botany and other branches of the natural history of the Himalayan Mountains, and of the flora of Cashmere. Wm. H. Allen & Co., London, 472 pp.
Schori, M. (2016) Cardiopteridaceae. In: Kadereit, J.W. & Bittrich, V. (Eds.) Flowering plants. Eudicots. The families and genera of vascular plants. Vol. 14. Springer, Cham, pp. 131–136. https://doi.org/10.1007/978-3-319-28534-4_8
Sleumer, H. (1969) Materials towards the knowledge of the Icacinaceae of Asia, Malesia, and adjacent areas. Blumea 17 (1): 181–264.
Sleumer, H. (1971) Icacinaceae. In: van Steenis, C.G.G.J. (Ed.) Flora Malesiana, series I, vol. 7 (1). Noordhoff, Leiden, pp. 1–87.
Stevens, P.F. (2001 onwards) Angiosperm Phylogeny Website. Version 14. Available from: http://www.mobot.org/MOBOT/research/APweb/ (accessed 22 October 2021)
Tagane, S., Wijedasa, L.S., Chhang, P., Toyama, H. & Yahara, T. (2015) Two new records for Cambodia’s forest flora, Memecylon corticosum var. kratense and M. paniculatum (Melastomataceae). Cambodian Journal of Natural History 2015 (2): 139–143.
Tagane, S., Toyama, H., Chhang, P., Dang, V.S. & Yahara, T. (2016a) New records of Xanthophyllum ellipticum and X. obscurum (Polygalaceae) in Indochina, with an identification key for species in the region. Cambodian Journal of Natural History 2016: 15–19.
Tagane, S., Naiki, A., Dang, V.S., Tran, H. & Yahara, T. (2016b) A new record of Gaertnera junghuhniana Miq. (Rubiaceae) from southern Vietnam. Acta Phytotaxonomica et Geobotanica 67 (2): 127–130. https://doi.org/10.18942/apg.201602
Tagane, S., Souladeth, P., Rueangruea, S., Okabe, N., Zhang, M., Chayer, S., Yang, C.J. & Yahara, T. (2018a) Flora of Nam Kading National Protected Area II: 30 new records of angiosperms to Laos. Edinburgh Journal of Botany 75 (1): 107–116. https://doi.org/10.1017/S0960428617000361
Tagane, S., Souladeth, P., Zhang, M. & Yahara, T. (2018b) Flora of Nam Kading National Protected Area IV: Two new species of Annonaceae, Monoon namkadingense and Neo-uvaria laosensis. Phytotaxa 336 (1): 82–88. http://dx.doi.org/10.11646/phytotaxa.336.1.6
Takhtajan, A.L. (2009) Flowering Plants. 2nd ed. Springer, New York, 871 pp. https://doi.org/10.1007/978-1-4020-9609-9
Truong, V.B., Truong, T.Q., Dang, V.S., Nguyen, T.M. & Vermeulen, J.J. (2020a) New records of Bulbophyllum in the Flora of Vietnam. Lankesteriana 20 (1): 31–36. http://dx.doi.org/10.15517/lank.v20i1.41130
Truong, V.B., Nguyen, V.C., Ormerod, P., Truong, Q.C., Chu, V.T. & Kumar, P. (2020b) The genus Corybas in Vietnam, with two new records. Lankesteriana 20 (1): 49–55. http://dx.doi.org/10.15517/lank.v20i1.41330
Utteridge, T.M.A. & Schori, M. (2011) Updating Malesian Icacinaceae. Gardens’ Bulletin Singapore 63 (1–2): 105–118.
Vislobokov, N.A., Nuraliev, M.S., Kuznetsov, A.N. & Kuznetsova, S.P. (2019) Aspidistra micrantha (Asparagaceae, Nolinoideae), a new species from Vietnam with unusually small flowers. Phytotaxa 422 (3): 289–294. https://doi.org/10.11646/phytotaxa.422.3.8
Wu, S.G., Xiang, J.Y., Phan, K.L. & Souliya, O. (2012) Some new records of ferns from Vietnam and Laos (3) Pteris (Pteridaceae). Plant Diversity & Resources 34 (1): 22–24. http://dx.doi.org/10.3724/sp.j.1143.2012.11100
Zhang, C., Zhang, T., Luebert, F., Xiang, Y., Huang, C.H., Hu, Y., Rees, M., Frohlich, M.W., Qi, J., Weigend, M. & Ma, H. (2020) Asterid phylogenomics/phylotranscriptomics uncover morphological evolutionary histories and support phylogenetic placement for numerous whole-genome duplications. Molecular Biology and Evolution 37 (11): 3188–3210. http://dx.doi.org/10.1093/molbev/msaa160