Abstract
Beauverdia hirtella comprises two subspecies: hirtella and lorentzii, the former occurs in Uruguay and southern Rio Grande do Sul state, Brazil, while the second subspecies spreads in southeastern Corrientes and eastern Entre Ríos provinces, Argentina. Based on field surveys, study of nomenclatural types and other specimens, and review of literature, the new subspecies Beauverdia hirtella subsp. glabrata is here proposed, being endemic to rocky grasslands in south-central Rio Grande do Sul state, Brazil. The new subspecies differs from the other two subspecies in several aspects, including the glabrous leaves and pedicels, the narrower leaves, and the smaller and nearly spherical or obovate capsules. Beauverdia hirtella subsp. glabrata is here described and illustrated, including comments on distribution, phenology and conservation. The main differences with regards to related taxa and a key to distinguish the taxa of Beauverdia are also provided.
References
<p>Baker, J.G. (1871) A Revision of the Genera and Species of Herbaceous Capsular Gamophyllous Liliaceae. <em>The Journal of the Linnean Society </em>11: 349–437. https://doi.org/10.1111/j.1095-8339.1870.tb00068.x</p>
<p>Baker, J.G. (1877) Systema Iridacearum. <em>Journal of Linnean Society, Botany</em> 16: 61–180. https://doi.org/10.1111/j.1095-8339.1877.tb00172.x</p>
<p>Baker, J.G. (1888) <em>Handbook of the Amaryllideae, including the Alstroemerieae and Agaveae</em>. George Bell & Sons, London, 216 pp. https://doi.org/10.5962/bhl.title.15516</p>
<p>Baker, J.G. (1892) <em>Handbook of the Irideae</em>. George Bells & Sons, London. 248 pp. https://doi.org/10.5962/bhl.title.15431</p>
<p>Baker, J.G. (1896) The genus <em>Brodiaea</em> and its allies. <em>The Gardeners’ chronicle: a weekly illustrated journal of horticulture and allied subjects</em>, ser. 3, 20: 459.</p>
<p>Beauverd, G. (1906) Una Liliacée nouvelle de l’Uruguay. <em>Bulletin de L´Herbier Boissier</em>, 2a. sér, 6: 1011.</p>
<p>Beauverd, G. (1908) Nouvelles espèces uruguayennes du genre <em>Nothoscordum</em> Kunth. <em>Bulletin de L´Herbier Boissier</em>, 2a. sér, 12: 994–1007.</p>
<p>Beauverd, G. (1921) Plantes Nouvelles de l’Uruguay. <em>Bulletin de la Société botanique de Genève</em>, 2a. sér., 13-14: 267–271.</p>
<p>Beentje, H. (2010) <em>The Kew Plant Glossary: an illustrated dictionary of plant terms</em>. Royal Botanic Gardens, Kew, 160 pp.</p>
<p>Bertero, C.L.G. (1829) Continuación del catálogo de plantas observadas en Chile. <em>El Mercurio Chileno </em>16: 735–749.</p>
<p>Bilenca, D. & Miñarro, F. (2004) <em>Identificación de áreas valiosas de pastizal en las Pampas y Campos de Argentina, Uruguay y Sur de Brasil</em>. Fundación Vida Silvestre, Buenos Aires, 353 pp.</p>
<p>Britton, N.L. & Brown, A. (1896) <em>An illustrated flora of the northern United States, Canada and the British possesions from Newfoundland to the parallel of the southern boundary of Virginia and from the Atlantic Ocean westward to the 102d meridian 1: Ophioglossaceae to Aizoaceae</em>. Charles Scriener’s Sons, New York, 612 pp. https://doi.org/10.5962/bhl.title.102081</p>
<p>Cabrera, Á.L. (1976) <em>Regiones Fitogeográficas Argentinas. Enciclopedia Argentina de Agricultura y Jardinería </em>Tomo II, Fascículo 1. Acme, Buenos Aires, 85 pp.</p>
<p>Cavanilles, A.J. (1793) <em>Icones et Descriptiones Plantarum</em> 2. Ex Regia typographia, Madrid, 83 pp, pl. 101–200.</p>
<p>Crosa, O. (1972) Estudios cariológicos en el género <em>Nothoscordum</em> (Liliaceae). <em>Boletín de la Facultad de Agronomía de la Universidad de Montevideo</em> 122: 3–8.</p>
<p>Crosa, O. (1975a) Las especies unifloras del género <em>Nothoscordum</em> Kunth y el género <em>Ipheion</em> Raf. de la tribu Allieae (Liliaceae). <em>Darwiniana</em> 19: 335–344.</p>
<p>Crosa, O. (1975b) <em>Zoellnerallium</em>, un género nuevo para la tribu Allieae (Liliaceae). <em>Darwiniana</em> 19: 331–334.</p>
<p>Crosa, O. (2005) Una nueva especie uniflora de <em>Nothoscordum</em> secc. <em>Nothoscordum</em> (Alliaceae) de Uruguay y sur de Brasil. <em>Hickenia </em>3: 253–256<em>.</em></p>
<p>Deble, L.P. (2011) Taxonomic novelties in <em>Calydorea</em> (Iridaceae: Tigridieae). <em>Bonplandia </em>20 (1): 35–39.</p>
<p>Garcia, N., Meerow, A., Arroyo-Leuenberg, S., Oliveira, R.S., Dutilh, J.H.A., Soltis, P.S. & Judd, W.S. (2019) Generic Classification of Amaryllidaceae tribe Hippeastreae. <em>Taxon </em>68 (3): 481–498. https://doi.org/10.1002/tax.12062</p>
<p>Goldblatt, P. (1978) <em>Herbertia</em> (Iridaceae) reinstated as a valid generic name. <em>Annals of the Missouri Botanic Garden</em> 64: 378–379. https://doi.org/10.2307/2395348</p>
<p>Grisebach, A. (1879) Symbolae ad floram Argentinam : Zweite Bearbeitung argentinischer Pflanzen nach den auf Befehl der H. Nacional-Regierung der argentinischen Republik durch die Professoren Lorentz und Hieronymus veranstalteten Sammlungen, sowie den im Museum zu Göttingen aufbewarhten Herbarien anderer Naturforscher, besonders den durch Herrn Schickendantz in der Provinz Catamarca gesammelten Pflanzen. <em>Abhandlungen der Königlichen Gesellschaft der Wissenschaften zu Göttingen</em> 24: 1–345. https://doi.org/10.5962/bhl.title.60503</p>
<p>Guaglianone, E.R. (1972) Sinopsis de las especies de <em>Ipheion</em> Raf. y <em>Nothoscordum</em> Kunth (Liliáceas) de Entre Ríos y regiones vecinas. <em>Darwiniana</em> 17: 159–242.</p>
<p>Herbert, W. (1821) <em>An Appendix: General index to the Botanical magazine</em>, <em>vol. 43-48 containing a treatise on bulbous roots with plates.</em> J. Ridgway and Sons, London, 52 pp.</p>
<p>Herbert, W. (1825) <em>Habranthus bifidus</em>: Two-cleft <em>Habranthus</em>. <em>Curtis’s Botanical Magazine</em> 52: pl. 2597.</p>
<p>Herbert, W. (1837) <em>Amaryllidaceae: preceded by an attempt to arrange the monocotyledonous orders, and followed by a treatise on cross-bred vegetables, and supplement</em>. J. Ridgway and Sons, London, 428 pp. https://doi.org/10.5962/bhl.title.116651</p>
<p>Herter, W.G. (1931) Florula Uruguayensis, Plantae vasculares<em>. Estudios Botánicos en la Región Uruguaya </em>4: 48.</p>
<p>Herter, W.G. (1943) <em>Beauverdia</em> genum novum Liliacearum. <em>Boissiera</em> 7: 505–512.</p>
<p>Herter, W.G. (1956) Flora del Uruguay. <em>Revista Sudamericana de Botánica</em> 1: 216.</p>
<p>Hieronymus, G. (1900) Selaginellarum species novae. I Species novae e sectione Homoeophyllarum Spring (Homotroparum Al. Br., subgeneris Euselaginellae Warb.), subsectione Rupestrium. <em>Hedwigia </em>39: 290–320.</p>
<p>IUCN Standards and Petitions Committee (2019) <em>Guidelines for Using the IUCN Red List Categories and Criteria</em>. Version 14. IUCN Species Survival Commission. IUCN, Gland, Switzerland and Cambridge, UK. [https://iucnredlist.org/redlist/content/attachmentfiles/RedListGuidelines.pdf]</p>
<p>Jackson, B.D. (1893) <em>Index Kewensis</em> 1: 689.</p>
<p>Kunth, K.S. (1843) Nothoscordum. <em>Enumeratio plantarum omnium hucusque cognitarum: secundum familias naturales disposita, adjectis characteribus, differentiis et synonymis </em>4: 457–464. https://doi.org/10.5962/bhl.title.67381</p>
<p>L’Heritier, C.L. (1788) <em>Sertum Anglicum: seu Plantae Rariores quae in Hortis juxta Londinum, Imprimis in Horto Regio Kewensi Excoluntur, ab Anno 1786 ad Annuum 1787 Observatae</em>. Typis Petri-Francisci Didot, Paris, 60 pp.</p>
<p>Lindley, J. (1830) <em>Triteleia. Edwards’s Botanical Register</em>15: 1217–1305.</p>
<p>Link, J.H.F. & Otto, C.F. (1828) <em>Ferraria pusilla. Icones plantarum selectarum</em> 10: 125–126.</p>
<p>Linnaeus, C. (1753) <em>Species plantarum :ex hibentes plantas rite cognitas, ad genera relatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas</em>. Holmiae, Impensis Laurentii Salvii, Stockholm, 560 pp. https://doi.org/10.5962/bhl.title.669</p>
<p>Molina, G.I. (1782) <em>Saggio sulla storia naturale del Chili</em>. Tipografia de’ Fratelli Masi e Comp., Bologna, 368 pp. https://doi.org/10.5962/bhl.title.62689</p>
<p>Molina, G.I. (1810) <em>Saggio sulla storia naturale del Chili</em>. 2ª. Tipografia de’ Fratelli Masi e Comp., Bologna, 309 pp. https://doi.org/10.5962/bhl.title.105203</p>
<p>Nees von Esenbeck, C.G.D. (1841) <em>Florae africae australioris illustrationes monographicae</em>. Prausnitzianis, Glogów, 490 pp.</p>
<p>Poepigg, E.F. (1833) <em>Fragmentum Synopseos Plantarum Phanerogamum ab Auctore Annis MDCCCXXVII ad MDCCCXXIX in Chile lectarum</em>. Lipseae [Leipzig], 33 pp.</p>
<p>Rafinesque, C.S (1836) <em>Flora Telluriana Pars Prima. Introd, et Classific. Ad mantissima synoptica 2000 Nova Genera plantarum vel nov. ord. et spec. In orbis Tellurianum 2</em>. H. Probasco, Philadelphia, 112 pp. https://doi.org/10.5962/bhl.title.7751</p>
<p>Ravenna, P.F. (1967a) <em>Nothoscordum felipponei</em> Beauv., its status and chromosomes. <em>Plant Life. Stanford, California </em>23: 49–52.</p>
<p>Ravenna, P.F. (1967b) Contribuições ao estudo das Amaryllidaceae da América do Sul. <em>Sellowia </em>19: 25–36.</p>
<p>Ravenna, P.F. (1968) <em>Nothoscordum</em> notes. <em>Plant Life. Stanford, California</em> 24: 57–58.</p>
<p>Ravenna, P.F. (1978) Studies in the Allieae II. <em>Plant Life. Stanford, California </em>34: 144.</p>
<p>Ravenna, P.F. (1981a) Eight new species and two new subspecies of <em>Cypella</em> (Iridaceae). <em>Wrightia</em> 7 (1): 13–21. https://doi.org/10.1111/j.1756-1051.1981.tb00714.x</p>
<p>Ravenna, P.F. (1981b) <em>Kelissa</em> a new genus of Iridaceae from South Brazil. <em>Adansonia</em> 1: 105–110.</p>
<p>Ravenna, P.F. (1983) <em>Catila</em> and <em>Onira</em>, two new genera of South America Iridaceae<em>. Nordic Journal of Botany</em> 3: 197–205. https://doi.org/10.1111/j.1756-1051.1983.tb01065.x</p>
<p>Ravenna, P.F. (1989) New species of <em>Nothoscordum </em>(Alliaceae) II. <em>Onira, Botanical Leaflets </em>2 (5): 41–43.</p>
<p>Ravenna, P.F. (1991) New species of <em>Nothoscordum </em>(Alliaceae) XIV. <em>Onira, Botanical Leaflets </em>3 (14): 40–41. https://doi.org/10.2307/1223230</p>
<p>Ravenna, P.F. (2001a) New or Noteworthy <em>Leucocoryne </em>species (Alliaceae) III. <em>Onira, Botanical Leaflets </em>5 (11): 42–43.</p>
<p>Ravenna, P.F. (2001b) New or critical <em>Tristagma</em> species (Alliaceae) I. <em>Onira, Botanical Leaflets</em> 6 (3): 24–33.</p>
<p>Rodrigues Souza, L.G., Crosa, O. & Guerra, M. (2010) Karyological circumscription of <em>Ipheion</em> Raf. (Gilliesioideae, Alliaceae). <em>Plant Systematics and Evolution</em> 28: 119–127. https://doi.org/10.1007/s00606-010-0304-3</p>
<p>Roitman, G. & Castillo, A. (2008) <em>Herbertia darwinii </em>(Iridaceae: Tigridieae: Cypurinae) a new species from South America. <em>Boletín Sociedad Argentina Botánica </em>43: 311–314.</p>
<p>Sassone, A.B., Giussani, L.M. & Guaglianone, E.R. (2013) Multivariate studies of <em>Ipheion</em> (Amaryllidaceae, Allioideae) and related genera. <em>Plant Systematics and Evolution</em> 299: 1561–1575. https://doi.org/10.1007/s00606-013-0819-5</p>
<p>Sassone, A.B, Giussani, L.M. & Guaglianone, E.R. (2014a) <em>Beauverdia</em>, a Resurrected Genus of Amaryllidaceae (Allioideae, Gilliesieae). <em>Systematic Botany </em>39 (3): 767–775. https://doi.org/10.1600/036364414X681527</p>
<p>Sassone, A.B., Arroyo-Leuenberger, S.C. & Giussani, L.M. (2014b) Nueva Circunscripción de la Tribu Leucocoryneae (Amaryllidaceae, Allioideae). <em>Darwiniana nueva serie </em>2: 197–206. https://doi.org/10.14522/darwiniana/2014.22.584</p>
<p>Sassone, A.B. & Giussani, L.M. (2018) Reconstructing the phylogenetic history of the tribe Leucocoryneae (Allioideae): Reticulate evolution and diversification in South America. <em>Molecular Phylogenetics and Evolution </em>127: 437–448. https://doi.org/10.1016/j.ympev.2018.04.034</p>
<p>Smith, J.E. (1810) Characters of a Liliaceous Genus called <em>Brodiaea</em>. <em>Transactions of the Linnean Society of London </em>10: 1–6. https://doi.org/10.1111/j.1096-3642.1810.tb00010.x</p>
<p>Sommer, C.A., Lima, E.F. de & Nardi, L.V.S. (1999) Evolução do vulcanismo alcalino na porção sul do Platô do Taquarembó, Dom Pedrito - RS. <em>Revista Brasileira de Geociências</em> 29 (2): 245–254. https://doi.org/10.25249/0375-7536.199929245254</p>
<p>Sommer, C.A., Lima, E.F., Pierosan, R. & Machado, A. (2011) Reoignimbritos e ignimbritos de alto grau do vulcanismo Acampamento Velho, RS: origem e temperatura de formação. <em>Revista Brasileira de Geociências </em>41 (3): 420–435. https://doi.org/10.25249/0375-7536.2011413420435</p>
<p>Soriano, A., León, R.J.C., Sala, O.E., Lavado, R.S., Deregibus, V.A., Cahuepé, O., Scaglia, A., Velazquez, C.A. & Lemcoff, J.H. (1992) Río de la Plata grasslands. <em>In</em>: Coupland, R.T. (Ed.) <em>Ecosystems of the World. Natural Grasslands. Introduction and Western Hemisphere</em>. Elsevier, Amsterdam, pp. 367–407.</p>
<p>Stearn, W.T. (1943) The Welsh onion and the ever-ready onion. <em>Gardeners’Chronicle</em> 1943: 60– 61. [London]</p>
<p>Sweet, R. (1829) <em>Herbertia</em> <em>pulchella</em>. Plaited leaved <em>Herbertia</em>. <em>The British flower garden: containing coloured figures & descriptions of the most ornamental & curious hardy herbaceous plants ... London</em> 3: pl. 222.</p>
<p>Traub, H.P. (1949) Amaryllid notes. <em>Plant Life. Stanford, California</em> 5: 50.</p>
<p>Traub, H.P. (1963) <em>Tristagma</em> Poepp. <em>Plant Life. Stanford, California</em> 19: 60–61.</p>
<p>Traub, H.P. & Moldenke, H.N. (1955) The genus <em>Ipheion</em>: Diagnosis, key to species and synonymy. <em>Plant Life. Stanford, California</em> 11: 125–130.</p>
<p>Thiers, B. (2021 [continuing database]) Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden´s Virtual Herbarium. Available from: http://sweetgum.nybg.org/ih/ (accessed 20 February 2021)</p>
<p>Wright, C.H. (1915) XV. – Decades Kewensis: Plantarum Novarum in Herbario Horti Regii Conservatarum. <em>Bulletin of Miscellaneous Information, Royal Gardens, Kew </em>1915 (3): 113–118. https://doi.org/10.2307/4107516</p>
<p>Zuccarini, J.G. (1831) Nachtrag zu der Monographie der amerikanischen Oxalis-Arten. <em>Abhandlungen der Mathematisch-Physikalischen Klasse der Königlich Bayerischen Akademie der Wissenschaften </em>1: 181–276.</p>
<p>Baker, J.G. (1877) Systema Iridacearum. <em>Journal of Linnean Society, Botany</em> 16: 61–180. https://doi.org/10.1111/j.1095-8339.1877.tb00172.x</p>
<p>Baker, J.G. (1888) <em>Handbook of the Amaryllideae, including the Alstroemerieae and Agaveae</em>. George Bell & Sons, London, 216 pp. https://doi.org/10.5962/bhl.title.15516</p>
<p>Baker, J.G. (1892) <em>Handbook of the Irideae</em>. George Bells & Sons, London. 248 pp. https://doi.org/10.5962/bhl.title.15431</p>
<p>Baker, J.G. (1896) The genus <em>Brodiaea</em> and its allies. <em>The Gardeners’ chronicle: a weekly illustrated journal of horticulture and allied subjects</em>, ser. 3, 20: 459.</p>
<p>Beauverd, G. (1906) Una Liliacée nouvelle de l’Uruguay. <em>Bulletin de L´Herbier Boissier</em>, 2a. sér, 6: 1011.</p>
<p>Beauverd, G. (1908) Nouvelles espèces uruguayennes du genre <em>Nothoscordum</em> Kunth. <em>Bulletin de L´Herbier Boissier</em>, 2a. sér, 12: 994–1007.</p>
<p>Beauverd, G. (1921) Plantes Nouvelles de l’Uruguay. <em>Bulletin de la Société botanique de Genève</em>, 2a. sér., 13-14: 267–271.</p>
<p>Beentje, H. (2010) <em>The Kew Plant Glossary: an illustrated dictionary of plant terms</em>. Royal Botanic Gardens, Kew, 160 pp.</p>
<p>Bertero, C.L.G. (1829) Continuación del catálogo de plantas observadas en Chile. <em>El Mercurio Chileno </em>16: 735–749.</p>
<p>Bilenca, D. & Miñarro, F. (2004) <em>Identificación de áreas valiosas de pastizal en las Pampas y Campos de Argentina, Uruguay y Sur de Brasil</em>. Fundación Vida Silvestre, Buenos Aires, 353 pp.</p>
<p>Britton, N.L. & Brown, A. (1896) <em>An illustrated flora of the northern United States, Canada and the British possesions from Newfoundland to the parallel of the southern boundary of Virginia and from the Atlantic Ocean westward to the 102d meridian 1: Ophioglossaceae to Aizoaceae</em>. Charles Scriener’s Sons, New York, 612 pp. https://doi.org/10.5962/bhl.title.102081</p>
<p>Cabrera, Á.L. (1976) <em>Regiones Fitogeográficas Argentinas. Enciclopedia Argentina de Agricultura y Jardinería </em>Tomo II, Fascículo 1. Acme, Buenos Aires, 85 pp.</p>
<p>Cavanilles, A.J. (1793) <em>Icones et Descriptiones Plantarum</em> 2. Ex Regia typographia, Madrid, 83 pp, pl. 101–200.</p>
<p>Crosa, O. (1972) Estudios cariológicos en el género <em>Nothoscordum</em> (Liliaceae). <em>Boletín de la Facultad de Agronomía de la Universidad de Montevideo</em> 122: 3–8.</p>
<p>Crosa, O. (1975a) Las especies unifloras del género <em>Nothoscordum</em> Kunth y el género <em>Ipheion</em> Raf. de la tribu Allieae (Liliaceae). <em>Darwiniana</em> 19: 335–344.</p>
<p>Crosa, O. (1975b) <em>Zoellnerallium</em>, un género nuevo para la tribu Allieae (Liliaceae). <em>Darwiniana</em> 19: 331–334.</p>
<p>Crosa, O. (2005) Una nueva especie uniflora de <em>Nothoscordum</em> secc. <em>Nothoscordum</em> (Alliaceae) de Uruguay y sur de Brasil. <em>Hickenia </em>3: 253–256<em>.</em></p>
<p>Deble, L.P. (2011) Taxonomic novelties in <em>Calydorea</em> (Iridaceae: Tigridieae). <em>Bonplandia </em>20 (1): 35–39.</p>
<p>Garcia, N., Meerow, A., Arroyo-Leuenberg, S., Oliveira, R.S., Dutilh, J.H.A., Soltis, P.S. & Judd, W.S. (2019) Generic Classification of Amaryllidaceae tribe Hippeastreae. <em>Taxon </em>68 (3): 481–498. https://doi.org/10.1002/tax.12062</p>
<p>Goldblatt, P. (1978) <em>Herbertia</em> (Iridaceae) reinstated as a valid generic name. <em>Annals of the Missouri Botanic Garden</em> 64: 378–379. https://doi.org/10.2307/2395348</p>
<p>Grisebach, A. (1879) Symbolae ad floram Argentinam : Zweite Bearbeitung argentinischer Pflanzen nach den auf Befehl der H. Nacional-Regierung der argentinischen Republik durch die Professoren Lorentz und Hieronymus veranstalteten Sammlungen, sowie den im Museum zu Göttingen aufbewarhten Herbarien anderer Naturforscher, besonders den durch Herrn Schickendantz in der Provinz Catamarca gesammelten Pflanzen. <em>Abhandlungen der Königlichen Gesellschaft der Wissenschaften zu Göttingen</em> 24: 1–345. https://doi.org/10.5962/bhl.title.60503</p>
<p>Guaglianone, E.R. (1972) Sinopsis de las especies de <em>Ipheion</em> Raf. y <em>Nothoscordum</em> Kunth (Liliáceas) de Entre Ríos y regiones vecinas. <em>Darwiniana</em> 17: 159–242.</p>
<p>Herbert, W. (1821) <em>An Appendix: General index to the Botanical magazine</em>, <em>vol. 43-48 containing a treatise on bulbous roots with plates.</em> J. Ridgway and Sons, London, 52 pp.</p>
<p>Herbert, W. (1825) <em>Habranthus bifidus</em>: Two-cleft <em>Habranthus</em>. <em>Curtis’s Botanical Magazine</em> 52: pl. 2597.</p>
<p>Herbert, W. (1837) <em>Amaryllidaceae: preceded by an attempt to arrange the monocotyledonous orders, and followed by a treatise on cross-bred vegetables, and supplement</em>. J. Ridgway and Sons, London, 428 pp. https://doi.org/10.5962/bhl.title.116651</p>
<p>Herter, W.G. (1931) Florula Uruguayensis, Plantae vasculares<em>. Estudios Botánicos en la Región Uruguaya </em>4: 48.</p>
<p>Herter, W.G. (1943) <em>Beauverdia</em> genum novum Liliacearum. <em>Boissiera</em> 7: 505–512.</p>
<p>Herter, W.G. (1956) Flora del Uruguay. <em>Revista Sudamericana de Botánica</em> 1: 216.</p>
<p>Hieronymus, G. (1900) Selaginellarum species novae. I Species novae e sectione Homoeophyllarum Spring (Homotroparum Al. Br., subgeneris Euselaginellae Warb.), subsectione Rupestrium. <em>Hedwigia </em>39: 290–320.</p>
<p>IUCN Standards and Petitions Committee (2019) <em>Guidelines for Using the IUCN Red List Categories and Criteria</em>. Version 14. IUCN Species Survival Commission. IUCN, Gland, Switzerland and Cambridge, UK. [https://iucnredlist.org/redlist/content/attachmentfiles/RedListGuidelines.pdf]</p>
<p>Jackson, B.D. (1893) <em>Index Kewensis</em> 1: 689.</p>
<p>Kunth, K.S. (1843) Nothoscordum. <em>Enumeratio plantarum omnium hucusque cognitarum: secundum familias naturales disposita, adjectis characteribus, differentiis et synonymis </em>4: 457–464. https://doi.org/10.5962/bhl.title.67381</p>
<p>L’Heritier, C.L. (1788) <em>Sertum Anglicum: seu Plantae Rariores quae in Hortis juxta Londinum, Imprimis in Horto Regio Kewensi Excoluntur, ab Anno 1786 ad Annuum 1787 Observatae</em>. Typis Petri-Francisci Didot, Paris, 60 pp.</p>
<p>Lindley, J. (1830) <em>Triteleia. Edwards’s Botanical Register</em>15: 1217–1305.</p>
<p>Link, J.H.F. & Otto, C.F. (1828) <em>Ferraria pusilla. Icones plantarum selectarum</em> 10: 125–126.</p>
<p>Linnaeus, C. (1753) <em>Species plantarum :ex hibentes plantas rite cognitas, ad genera relatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas</em>. Holmiae, Impensis Laurentii Salvii, Stockholm, 560 pp. https://doi.org/10.5962/bhl.title.669</p>
<p>Molina, G.I. (1782) <em>Saggio sulla storia naturale del Chili</em>. Tipografia de’ Fratelli Masi e Comp., Bologna, 368 pp. https://doi.org/10.5962/bhl.title.62689</p>
<p>Molina, G.I. (1810) <em>Saggio sulla storia naturale del Chili</em>. 2ª. Tipografia de’ Fratelli Masi e Comp., Bologna, 309 pp. https://doi.org/10.5962/bhl.title.105203</p>
<p>Nees von Esenbeck, C.G.D. (1841) <em>Florae africae australioris illustrationes monographicae</em>. Prausnitzianis, Glogów, 490 pp.</p>
<p>Poepigg, E.F. (1833) <em>Fragmentum Synopseos Plantarum Phanerogamum ab Auctore Annis MDCCCXXVII ad MDCCCXXIX in Chile lectarum</em>. Lipseae [Leipzig], 33 pp.</p>
<p>Rafinesque, C.S (1836) <em>Flora Telluriana Pars Prima. Introd, et Classific. Ad mantissima synoptica 2000 Nova Genera plantarum vel nov. ord. et spec. In orbis Tellurianum 2</em>. H. Probasco, Philadelphia, 112 pp. https://doi.org/10.5962/bhl.title.7751</p>
<p>Ravenna, P.F. (1967a) <em>Nothoscordum felipponei</em> Beauv., its status and chromosomes. <em>Plant Life. Stanford, California </em>23: 49–52.</p>
<p>Ravenna, P.F. (1967b) Contribuições ao estudo das Amaryllidaceae da América do Sul. <em>Sellowia </em>19: 25–36.</p>
<p>Ravenna, P.F. (1968) <em>Nothoscordum</em> notes. <em>Plant Life. Stanford, California</em> 24: 57–58.</p>
<p>Ravenna, P.F. (1978) Studies in the Allieae II. <em>Plant Life. Stanford, California </em>34: 144.</p>
<p>Ravenna, P.F. (1981a) Eight new species and two new subspecies of <em>Cypella</em> (Iridaceae). <em>Wrightia</em> 7 (1): 13–21. https://doi.org/10.1111/j.1756-1051.1981.tb00714.x</p>
<p>Ravenna, P.F. (1981b) <em>Kelissa</em> a new genus of Iridaceae from South Brazil. <em>Adansonia</em> 1: 105–110.</p>
<p>Ravenna, P.F. (1983) <em>Catila</em> and <em>Onira</em>, two new genera of South America Iridaceae<em>. Nordic Journal of Botany</em> 3: 197–205. https://doi.org/10.1111/j.1756-1051.1983.tb01065.x</p>
<p>Ravenna, P.F. (1989) New species of <em>Nothoscordum </em>(Alliaceae) II. <em>Onira, Botanical Leaflets </em>2 (5): 41–43.</p>
<p>Ravenna, P.F. (1991) New species of <em>Nothoscordum </em>(Alliaceae) XIV. <em>Onira, Botanical Leaflets </em>3 (14): 40–41. https://doi.org/10.2307/1223230</p>
<p>Ravenna, P.F. (2001a) New or Noteworthy <em>Leucocoryne </em>species (Alliaceae) III. <em>Onira, Botanical Leaflets </em>5 (11): 42–43.</p>
<p>Ravenna, P.F. (2001b) New or critical <em>Tristagma</em> species (Alliaceae) I. <em>Onira, Botanical Leaflets</em> 6 (3): 24–33.</p>
<p>Rodrigues Souza, L.G., Crosa, O. & Guerra, M. (2010) Karyological circumscription of <em>Ipheion</em> Raf. (Gilliesioideae, Alliaceae). <em>Plant Systematics and Evolution</em> 28: 119–127. https://doi.org/10.1007/s00606-010-0304-3</p>
<p>Roitman, G. & Castillo, A. (2008) <em>Herbertia darwinii </em>(Iridaceae: Tigridieae: Cypurinae) a new species from South America. <em>Boletín Sociedad Argentina Botánica </em>43: 311–314.</p>
<p>Sassone, A.B., Giussani, L.M. & Guaglianone, E.R. (2013) Multivariate studies of <em>Ipheion</em> (Amaryllidaceae, Allioideae) and related genera. <em>Plant Systematics and Evolution</em> 299: 1561–1575. https://doi.org/10.1007/s00606-013-0819-5</p>
<p>Sassone, A.B, Giussani, L.M. & Guaglianone, E.R. (2014a) <em>Beauverdia</em>, a Resurrected Genus of Amaryllidaceae (Allioideae, Gilliesieae). <em>Systematic Botany </em>39 (3): 767–775. https://doi.org/10.1600/036364414X681527</p>
<p>Sassone, A.B., Arroyo-Leuenberger, S.C. & Giussani, L.M. (2014b) Nueva Circunscripción de la Tribu Leucocoryneae (Amaryllidaceae, Allioideae). <em>Darwiniana nueva serie </em>2: 197–206. https://doi.org/10.14522/darwiniana/2014.22.584</p>
<p>Sassone, A.B. & Giussani, L.M. (2018) Reconstructing the phylogenetic history of the tribe Leucocoryneae (Allioideae): Reticulate evolution and diversification in South America. <em>Molecular Phylogenetics and Evolution </em>127: 437–448. https://doi.org/10.1016/j.ympev.2018.04.034</p>
<p>Smith, J.E. (1810) Characters of a Liliaceous Genus called <em>Brodiaea</em>. <em>Transactions of the Linnean Society of London </em>10: 1–6. https://doi.org/10.1111/j.1096-3642.1810.tb00010.x</p>
<p>Sommer, C.A., Lima, E.F. de & Nardi, L.V.S. (1999) Evolução do vulcanismo alcalino na porção sul do Platô do Taquarembó, Dom Pedrito - RS. <em>Revista Brasileira de Geociências</em> 29 (2): 245–254. https://doi.org/10.25249/0375-7536.199929245254</p>
<p>Sommer, C.A., Lima, E.F., Pierosan, R. & Machado, A. (2011) Reoignimbritos e ignimbritos de alto grau do vulcanismo Acampamento Velho, RS: origem e temperatura de formação. <em>Revista Brasileira de Geociências </em>41 (3): 420–435. https://doi.org/10.25249/0375-7536.2011413420435</p>
<p>Soriano, A., León, R.J.C., Sala, O.E., Lavado, R.S., Deregibus, V.A., Cahuepé, O., Scaglia, A., Velazquez, C.A. & Lemcoff, J.H. (1992) Río de la Plata grasslands. <em>In</em>: Coupland, R.T. (Ed.) <em>Ecosystems of the World. Natural Grasslands. Introduction and Western Hemisphere</em>. Elsevier, Amsterdam, pp. 367–407.</p>
<p>Stearn, W.T. (1943) The Welsh onion and the ever-ready onion. <em>Gardeners’Chronicle</em> 1943: 60– 61. [London]</p>
<p>Sweet, R. (1829) <em>Herbertia</em> <em>pulchella</em>. Plaited leaved <em>Herbertia</em>. <em>The British flower garden: containing coloured figures & descriptions of the most ornamental & curious hardy herbaceous plants ... London</em> 3: pl. 222.</p>
<p>Traub, H.P. (1949) Amaryllid notes. <em>Plant Life. Stanford, California</em> 5: 50.</p>
<p>Traub, H.P. (1963) <em>Tristagma</em> Poepp. <em>Plant Life. Stanford, California</em> 19: 60–61.</p>
<p>Traub, H.P. & Moldenke, H.N. (1955) The genus <em>Ipheion</em>: Diagnosis, key to species and synonymy. <em>Plant Life. Stanford, California</em> 11: 125–130.</p>
<p>Thiers, B. (2021 [continuing database]) Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden´s Virtual Herbarium. Available from: http://sweetgum.nybg.org/ih/ (accessed 20 February 2021)</p>
<p>Wright, C.H. (1915) XV. – Decades Kewensis: Plantarum Novarum in Herbario Horti Regii Conservatarum. <em>Bulletin of Miscellaneous Information, Royal Gardens, Kew </em>1915 (3): 113–118. https://doi.org/10.2307/4107516</p>
<p>Zuccarini, J.G. (1831) Nachtrag zu der Monographie der amerikanischen Oxalis-Arten. <em>Abhandlungen der Mathematisch-Physikalischen Klasse der Königlich Bayerischen Akademie der Wissenschaften </em>1: 181–276.</p>