Abstract
Asplenium pahli, a new species from Leyte, Philippines, is described. Morphological and distribution range differences between A. pahli, A. anisodontum, A. caudatum, A. falcatum, A. gueinzianum, and A. planicaule are discussed. The habitat and climate of Eastern Leyte is described.
References
<p>Blume, C.L. (1828) <em>Enumeratio Plantarum Javae et insularum adjacentium</em>. J.W. Van Leeuwen, Lugduni Batavorum (Leiden), 247 pp. https://doi.org/10.5962/bhl.title.44901</p>
<p>Burman, J. (1737) <em>Thesaurus Zeylanicus, exhibens Plantas in Insula Zeylania Nascentes</em>. Janssonio- Waesbergios, Salomonem Schouten, Amstelaedami, 237 pp https://doi.org/10.5962/bhl.title.734</p>
<p>Forster, G. (1786) <em>Florulae Insularum Australium Prodromus</em>. Joannes Christian Dietrich, Göttingen, Germany, 103 pp. https://doi.org/10.5962/bhl.title.10725</p>
<p>Fraser-Jenkins, C.R. (1992) The ferns and fern-allies of the far west Himalaya. <em>Pakistan Systematics</em> 5: 85–120.</p>
<p>Goode, J.P. (1912) The Forest of the Philippines. <em>Bulletin of the American Geographical Society</em> 44: 81–89. https://doi.org/10.2307/200216</p>
<p>Kuhn, M. (1868) <em>Filices Africanae</em>. W. Engelmann, Lipsiae. 233 pp.</p>
<p>Kunze, G. (1846) Filices Javae Zollingerianas aliasque ex herbario Moricandiano observationes Kunzii, Lipsiensis. <em>Botanische Zeitung</em> 4: 44−447. [https://www.biodiversitylibrary.org/item/101480#page/233/mode/1up]</p>
<p>Lamarck, J.B.P. (1786) <em>Encyclopédie Méthodique Botanique 2</em>. Panckoucke, Plomteux, Paris, Liege, 774 pp. https://doi.org/10.5962/bhl.title.824</p>
<p>Lasco, R.D., Visco, R.G. & Pulhin, J.M. (2001) Secondary forests in the Philippines: Formation and transformation in the 20th century. <em>Journal of Tropical Forest Science</em> 13: 652−670. Available from: https://www.researchgate.net/publication/239592580_Secondary_forests_in_the_Philippines_Formation_and_transformation_in_the_20th_century (accessed 3 November 2020)</p>
<p>Lellinger, D.B. (2002) <em>A modern multilingual glossary for taxonomic pteridology</em>. Pteridologia 3, American Fern Society, Washington, DC, United States, 263 pp. https://doi.org/10.5962/bhl.title.124209</p>
<p>Linnaeus, C. (1753) <em>Species Plantarum, exhibentes plantas rite cognitas, ad genera relates cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum system sexuale digestas</em>, vol. 2. Laurentii Salvii, Holmiae, 970 pp. https://doi.org/10.5962/bhl.title.669</p>
<p>Makino, T. (1900) <em>Shinsen Nihon shokubutsu zusetsu</em> [Phanerogamae et Pteridophytae Japonicae Iconibus Illustratae] 2: pl. 64. Keigyosha, Meiji 32−36, Tokyo. Available from: https://www.biodiversitylibrary.org/item/111978#page/348/mode/1up (accessed 26 November 2020).</p>
<p>Mettenius, G.H. (1859) Ueber einige Farngattungen. Asplenium.<em> In: Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft </em>vol. 3. Heinrich Ludwig Broenner, Frankfurt am Main. pp. 100−254. Available from: https://www.biodiversitylibrary.org/pdf4/122160900188342.pdf (accessed 27 November 2020)</p>
<p>Mongabay (2020) <em>Deforestation statistics for the Philippines</em>. Available from: https://rainforests.mongabay.com/deforestation/archive/Philippines.htm (accessed 8 September 2020)</p>
<p>Morton, C. (1967) Studies of fern types. I. <em>Contributions from the United States National Herbarium</em> 38: 29− 83. Available from: http://www.jstor.org/stable/23493033 (accessed 8 September 2020)</p>
<p>Morton, C. (1973) Studies of fern types. II. <em>Contributions from the United States National Herbarium</em> 38: 215−229. Available from: https://www.jstor.org/stable/23493107 (accessed 29 November 2020)</p>
<p>Nicolson DH, Fosberg FR. (2003) The Forsters and the botany of the second Cook expedition (1772–1775). <em>Regnum Vegetabile</em> 139: 1–760.</p>
<p>Pagasa. (2020) Philippine Atmospheric, Geophysical and Astronomical Services Administration (PAGASA). Available from: http://bagong.pagasa.dost.gov.ph/climate/climatological-normals. (accessed 7 September 2020)</p>
<p>Presl, C. (1849) <em>Epimeliae Botanicae: cum tabulis quindecim lithographicis</em>. Amadei Haase, Pragae, 264 pp. https://doi.org/10.5962/bhl.title.61845</p>
<p>Salgado, A.E. (1996) Taxonomic and Nomenclatural Notes on Philippine Ferns. I. On the Identity of <em>Asplenium anisodontum</em> C.Presl. <em>Edinburgh Journal of Botany</em> 53: 271−274. https://doi.org/10.1017/S0960428600002900</p>
<p>Salgado, A.E. (2017) Does <em>Asplenium falcatum</em> exist in the Philippines? <em>Edinburgh Journal of Botany</em> 74: 1− 4. https://doi.org/10.1017/S0960428616000214</p>
<p>Salgado, A.E. (2020) Taxonomic and Nomenclatural Notes on Philippine Ferns. III. <em>Asplenium caudatum</em> G.Forst. (Aspleniaceae) and ferns confused with it. <em>Fern Gazette</em> 21: 169−184.</p>
<p>Salgado, A.E. & C.R. Fraser-Jenkins (2013) The nomenclature, typification and taxonomy of <em>Asplenium</em> <em>falcatum</em>, <em>A. polyodon</em> and confused species. <em>Fern Gazette</em> 19: 213−239. [https://ebps.org.uk/wp-content/uploads/2014/06/FGV19P6M2-42qQdr.pdf]</p>
<p>Sledge, W.A. (1965) The Ceylon Species of <em>Asplenium</em>. <em>Bulletin of the British Museum (Natural History). Botany</em> 3: 233−286.</p>
<p>Steenis-Kruseman, M.J. van. (1950) Alphabetical List of Collectors.<em> In: </em>Steenis, C.G.G.J. van (ed.) <em>Flora Malesiana. Series I, Spermatophyta</em>, vol. 1. Special Part. Departemen Pertanian, Kebun Raya, Lembaga Ilmu Pengetahuan, Noordhoff-Kolff N.V., Djakarta, Indonesia, 606 pp. https://doi.org/10.5962/bhl.title.40744</p>
<p>Swartz, O. (1800) Genera et Species Filicum ordine systematico redactarum. <em>Schrader Journal of Botany</em> 2: 1−120.</p>
<p>Wallich, N. (1828) <em>Numerical list of dried specimens of plants in the Museum of the Honl. East India Company</em>. Available from: https://www.biodiversitylibrary.org/item/15503#page/1/mode/1up (accessed 26 November 2020). https://doi.org/10.5962/bhl.title.1917</p>
<p>Wenzel, C.A. (1913) 16 March 1913. Letter of C.A. Wenzel to Sir David Prain from Jaro, Leyte, Philippines. Royal Botanic Gardens, Kew. Directors’ Correspondence, vol. 167, folio 391. Library and Archives at the Royal Botanic Gardens, Kew, England. Available <br>from: https://plants.jstor.org/stable/10.5555/al.ap.visual.kdcas9949 (accessed 2 September 2021)</p>
<p>Burman, J. (1737) <em>Thesaurus Zeylanicus, exhibens Plantas in Insula Zeylania Nascentes</em>. Janssonio- Waesbergios, Salomonem Schouten, Amstelaedami, 237 pp https://doi.org/10.5962/bhl.title.734</p>
<p>Forster, G. (1786) <em>Florulae Insularum Australium Prodromus</em>. Joannes Christian Dietrich, Göttingen, Germany, 103 pp. https://doi.org/10.5962/bhl.title.10725</p>
<p>Fraser-Jenkins, C.R. (1992) The ferns and fern-allies of the far west Himalaya. <em>Pakistan Systematics</em> 5: 85–120.</p>
<p>Goode, J.P. (1912) The Forest of the Philippines. <em>Bulletin of the American Geographical Society</em> 44: 81–89. https://doi.org/10.2307/200216</p>
<p>Kuhn, M. (1868) <em>Filices Africanae</em>. W. Engelmann, Lipsiae. 233 pp.</p>
<p>Kunze, G. (1846) Filices Javae Zollingerianas aliasque ex herbario Moricandiano observationes Kunzii, Lipsiensis. <em>Botanische Zeitung</em> 4: 44−447. [https://www.biodiversitylibrary.org/item/101480#page/233/mode/1up]</p>
<p>Lamarck, J.B.P. (1786) <em>Encyclopédie Méthodique Botanique 2</em>. Panckoucke, Plomteux, Paris, Liege, 774 pp. https://doi.org/10.5962/bhl.title.824</p>
<p>Lasco, R.D., Visco, R.G. & Pulhin, J.M. (2001) Secondary forests in the Philippines: Formation and transformation in the 20th century. <em>Journal of Tropical Forest Science</em> 13: 652−670. Available from: https://www.researchgate.net/publication/239592580_Secondary_forests_in_the_Philippines_Formation_and_transformation_in_the_20th_century (accessed 3 November 2020)</p>
<p>Lellinger, D.B. (2002) <em>A modern multilingual glossary for taxonomic pteridology</em>. Pteridologia 3, American Fern Society, Washington, DC, United States, 263 pp. https://doi.org/10.5962/bhl.title.124209</p>
<p>Linnaeus, C. (1753) <em>Species Plantarum, exhibentes plantas rite cognitas, ad genera relates cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum system sexuale digestas</em>, vol. 2. Laurentii Salvii, Holmiae, 970 pp. https://doi.org/10.5962/bhl.title.669</p>
<p>Makino, T. (1900) <em>Shinsen Nihon shokubutsu zusetsu</em> [Phanerogamae et Pteridophytae Japonicae Iconibus Illustratae] 2: pl. 64. Keigyosha, Meiji 32−36, Tokyo. Available from: https://www.biodiversitylibrary.org/item/111978#page/348/mode/1up (accessed 26 November 2020).</p>
<p>Mettenius, G.H. (1859) Ueber einige Farngattungen. Asplenium.<em> In: Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft </em>vol. 3. Heinrich Ludwig Broenner, Frankfurt am Main. pp. 100−254. Available from: https://www.biodiversitylibrary.org/pdf4/122160900188342.pdf (accessed 27 November 2020)</p>
<p>Mongabay (2020) <em>Deforestation statistics for the Philippines</em>. Available from: https://rainforests.mongabay.com/deforestation/archive/Philippines.htm (accessed 8 September 2020)</p>
<p>Morton, C. (1967) Studies of fern types. I. <em>Contributions from the United States National Herbarium</em> 38: 29− 83. Available from: http://www.jstor.org/stable/23493033 (accessed 8 September 2020)</p>
<p>Morton, C. (1973) Studies of fern types. II. <em>Contributions from the United States National Herbarium</em> 38: 215−229. Available from: https://www.jstor.org/stable/23493107 (accessed 29 November 2020)</p>
<p>Nicolson DH, Fosberg FR. (2003) The Forsters and the botany of the second Cook expedition (1772–1775). <em>Regnum Vegetabile</em> 139: 1–760.</p>
<p>Pagasa. (2020) Philippine Atmospheric, Geophysical and Astronomical Services Administration (PAGASA). Available from: http://bagong.pagasa.dost.gov.ph/climate/climatological-normals. (accessed 7 September 2020)</p>
<p>Presl, C. (1849) <em>Epimeliae Botanicae: cum tabulis quindecim lithographicis</em>. Amadei Haase, Pragae, 264 pp. https://doi.org/10.5962/bhl.title.61845</p>
<p>Salgado, A.E. (1996) Taxonomic and Nomenclatural Notes on Philippine Ferns. I. On the Identity of <em>Asplenium anisodontum</em> C.Presl. <em>Edinburgh Journal of Botany</em> 53: 271−274. https://doi.org/10.1017/S0960428600002900</p>
<p>Salgado, A.E. (2017) Does <em>Asplenium falcatum</em> exist in the Philippines? <em>Edinburgh Journal of Botany</em> 74: 1− 4. https://doi.org/10.1017/S0960428616000214</p>
<p>Salgado, A.E. (2020) Taxonomic and Nomenclatural Notes on Philippine Ferns. III. <em>Asplenium caudatum</em> G.Forst. (Aspleniaceae) and ferns confused with it. <em>Fern Gazette</em> 21: 169−184.</p>
<p>Salgado, A.E. & C.R. Fraser-Jenkins (2013) The nomenclature, typification and taxonomy of <em>Asplenium</em> <em>falcatum</em>, <em>A. polyodon</em> and confused species. <em>Fern Gazette</em> 19: 213−239. [https://ebps.org.uk/wp-content/uploads/2014/06/FGV19P6M2-42qQdr.pdf]</p>
<p>Sledge, W.A. (1965) The Ceylon Species of <em>Asplenium</em>. <em>Bulletin of the British Museum (Natural History). Botany</em> 3: 233−286.</p>
<p>Steenis-Kruseman, M.J. van. (1950) Alphabetical List of Collectors.<em> In: </em>Steenis, C.G.G.J. van (ed.) <em>Flora Malesiana. Series I, Spermatophyta</em>, vol. 1. Special Part. Departemen Pertanian, Kebun Raya, Lembaga Ilmu Pengetahuan, Noordhoff-Kolff N.V., Djakarta, Indonesia, 606 pp. https://doi.org/10.5962/bhl.title.40744</p>
<p>Swartz, O. (1800) Genera et Species Filicum ordine systematico redactarum. <em>Schrader Journal of Botany</em> 2: 1−120.</p>
<p>Wallich, N. (1828) <em>Numerical list of dried specimens of plants in the Museum of the Honl. East India Company</em>. Available from: https://www.biodiversitylibrary.org/item/15503#page/1/mode/1up (accessed 26 November 2020). https://doi.org/10.5962/bhl.title.1917</p>
<p>Wenzel, C.A. (1913) 16 March 1913. Letter of C.A. Wenzel to Sir David Prain from Jaro, Leyte, Philippines. Royal Botanic Gardens, Kew. Directors’ Correspondence, vol. 167, folio 391. Library and Archives at the Royal Botanic Gardens, Kew, England. Available <br>from: https://plants.jstor.org/stable/10.5555/al.ap.visual.kdcas9949 (accessed 2 September 2021)</p>