Abstract
Tribe Vaccinieae Reichenbach with more than 800 species contains most of the neotropical taxa of Ericaceae Jussieu (Pedraza-Peñalosa 2015a: 35). Species of genus Vaccinium Linneaus (1753: 349) known as blueberries or mortiños in Spanish, are members of Vaccinieae well-known for their economic importance (Pedraza-Peñalosa 2015b: 100). The genus Ceratostema Jussieu (1789: 163) was created by Antoine Jussieu and Ceratostema peruvianum Gmelin (1791: 676) was designated as the type species (Luteyn 1986: 485). It was first treated by Smith in 1952, who published a brief description and an artificial key, recognizing 16 species at that time (Smith 1952). Luteyn (1986) reviewed the genus and provided an updated artificial key to 23 species, including three new ones. Subsequently, a new species from Peru was added and thirteen more from Ecuador were included in the genus (Luteyn 1992, 1996, 2005).
References
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<p>Pedraza-Peñalosa, P. (2015b) <em>Themistoclesia diazii</em>, una especie nueva relacionada con los mortiños colombianos (Ericaceae: Vaccinieae). <em>Caldasia</em> 37: 99–100. https://doi.org/10.15446/caldasia.v37n1.50817</p>
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<p>Sleumer, H.O. (1941) Vaccinioideen-Studien. <em>Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie</em> 71: 375–510.</p>
<p>Smith, A.C. (1952) Plants collected in Ecuador by W. H. Camp. Vaccineae. <em>Memoirs of the New York Botanical Garden </em>8: 58–59.</p>
<p>Willdenow, C.L. von (1806) <em>Species plantarum</em>, Tomus IV. Pars II. Laurentius Salvius, Stockholm, pp. 633–1157.</p>
<p>Britton, N.L. & Rose, J.N. (1928) <em>North American Flora.</em> <em>Volume 23. Part I. (Rosales). Mimosaceae</em>. The New York Botanical Garden, New York City, 76 pp.</p>
<p>Gerique, A. (2011) <em>Biodiversity as a resource: plant use and land use among the Shuar, Saraguros, and Mestizos in tropical rainforest areas of southern Ecuador. </em>Friedrich-Alexander-Universität Erlangen-Nuremberg, 429 pp.</p>
<p>Gmelin, J.F. (1791) <em>Caroli a Linné systema naturae per regna tria naturae. Tomus II. Editio decima tertia, aucta, reformata</em> <em>2 parts</em>. Leipzig. (Syst. Nat.), 884 pp.</p>
<p>Hoerold, R. (1909) Systematische Gliederung und geographische Verbreitung der amerikanischen Thibaudieen. <em>Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie</em> 13: 251–334.</p>
<p>Hooker, W.J. (1837) <em>Macleania floribunda.</em> <em>Icones Plantarum</em> 2: t. 109.</p>
<p>IUCN. (2019) <em>Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. </em>Prepared by the Standards and Petitions Subcommittee in August 2019. Available from: https://cmsdocs.s3.amazonaws.com/RedListGuidelines.pdf (accessed 13 August 2021)</p>
<p>Jørgensen, P.M. & León-Yánez, S. (1999) Catálogo de las plantas vasculares del Ecuador. <em>Monographs in Systematic Botany from the Missouri Botanical Garden </em>75: 1–1182.</p>
<p>Jussieu, A.L. de (1789) <em>Genera plantarum secundum ordines naturales disposita, juxta methoclum in horto regio parisiensi exaratum</em>. Viduam Herissant and Theophilum Barrois, Paris, 498 pp. https://doi.org/10.5962/bhl.title.7762</p>
<p>Lind, R. (2012) <em>Open source software for image processing and analysis: picture this with ImageJ. In: </em>Harland, L. & Forster, M. (Eds.)<em> Open Source Software in Life Science Research: Practical Solutions in the Pharmaceutical Industry and Beyond: </em>Abington, Cambridge UK, Woodhead Publishing Ltd., pp. 131–149. https://doi.org/10.1533/9781908818249.131</p>
<p>Linnaeus, C. (1753) <em>Species plantarum</em>, Tomus I. Laurentius Salvius, Stockholm, 560 pp.</p>
<p>Lozano, P., Bussmann, R.W. & Küppers, M. (2007) Diversidad florística del bosque montano en el Occidente del Parque Nacional Podocarpus, Sur del Ecuador y su influencia en la flora pionera en deslizamientos naturales. <em>Revista UDO Agrícola</em> 7: 142–159. </p>
<p>Luteyn, J.L. (1986) New species of <em>Ceratostema</em> (Ericaceae: Vaccineae) from the northern Andes. <em>Journal of the Arnold Arboretum</em> 67: 485–492. https://doi.org/10.5962/bhl.part.27395</p>
<p>Luteyn, J.L. (1992) A New <em>Ceratostema</em> (Ericaceae: Vaccinieae) from Northern Peru. <em>Bulletin of the Torrey Botanical Club</em> 119: 314–315. https://doi.org/10.2307/2996763</p>
<p>Luteyn, J.L. (1996) Ericaceae: <em>Ceratostema.</em> <em>In: </em>Harling, G. & Andersson, L. (Eds.) <em>Flora of Ecuador. Volume 54. Part 147</em>. Göteborg University, Stockholm, pp. 210–253.</p>
<p>Luteyn, J.L. (2005) Four new species of Ericaceae (Vaccinieae) from Ecuador.<em> SIDA, Contributions to Botany </em>21: 1269–1282. </p>
<p>Pedraza-Peñalosa, P. (2015a) New blueberry and mortiño relatives (Ericaceae) from northwestern Colombia. <em>PhytoKeys</em> 49: 33–58 https://doi.org/10.3897/phytokeys.49.8383</p>
<p>Pedraza-Peñalosa, P. (2015b) <em>Themistoclesia diazii</em>, una especie nueva relacionada con los mortiños colombianos (Ericaceae: Vaccinieae). <em>Caldasia</em> 37: 99–100. https://doi.org/10.15446/caldasia.v37n1.50817</p>
<p>Sleumer, H.O. (1935) Ericaceae americanae etc. II. Beiträge zur Kenntnis der Flacourtiaceen Südamerikas II. <em>Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem</em> 12: 277–294. https://doi.org/10.2307/3994889</p>
<p>Sleumer, H.O. (1941) Vaccinioideen-Studien. <em>Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie</em> 71: 375–510.</p>
<p>Smith, A.C. (1952) Plants collected in Ecuador by W. H. Camp. Vaccineae. <em>Memoirs of the New York Botanical Garden </em>8: 58–59.</p>
<p>Willdenow, C.L. von (1806) <em>Species plantarum</em>, Tomus IV. Pars II. Laurentius Salvius, Stockholm, pp. 633–1157.</p>