Abstract
The nomenclature that Perrier de la Bâthie and Boiteau & Mannoni successively published for the Malagasy K. miniata (Crassulaceae subfam Kalanchooideae) in the 1920s and 1940s, respectively, is analysed. New synonyms are recorded for K. miniata and designations not validly published are noted. The infraspecific names published in K. miniata by Perrier de la Bâthie and Boiteau & Mannoni are interpreted as referring to material that belong to three species, namely to K. miniata, K. pseudocampanulata, and K. pubescens. Several names are typified and a new combination, K. pubescens var. tsinjoarivensis, based on K. miniata var. tsinjoarivensis, is published.
References
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Baker, J.G. (1881 [21 February]) Notes on a collection of flowering plants made by L. Kitching, Esq., in Madagascar in 1879 (Plates VII. & VIII.). Journal of the Linnean Society, Botany 18: 264–281. https://doi.org/10.1111/j.1095-8339.1881.tb01257.x
Baker, J.G. (1882) Contributions to the flora of central Madagascar (to be continued). Journal of Botany, British and Foreign. London 20: 109–114. [https://www.biodiversitylibrary.org/item/35873#page/125/mode/1up]
Baker, J.G. (1887) Further contributions to the flora of Madagascar. Journal of the Linnean Society, Botany (London) 22: 441–537. https://doi.org/10.1111/j.1095-8339.1887.tb00472.x
Berger, A. (1930 [Mai]) Crassulaceae. Unterfam. II. Kalanchoideae [sic, Kalanchooideae]. 6. Kalanchoe Adanson. 7. Kitchingia Baker. 8. Bryophyllum Salisb. In: A. Engler & K. Prantl (Eds.) Die natürlichen Pflanzenfamilien. ed. 2, 18a. Verlag von Wilhelm Engelmann, Leipzig, pp. 402–412.
Boiteau, P. & Allorge-Boiteau, L. (1995). Kalanchoe (Crassulacées) de Madagascar. Systématique, écophysiologie et phytochimie. ICSN, CNRS, 91198 Gif-sur-Yvette, Éditions Karthala, Paris, 252 pp.
Boiteau, P. & Mannoni, O. (1948a [mars-juin]). Les Kalanchoe (suite). Section II. Bryophyllum. [§ 3. Centrales; Kalanchoe pseudocampanulata Mannoni & Boiteau; K. rolandi-bonapartei Raym.-Hamet & H.Perrier; K. pubescens Baker] (à suivre). Cactus (Paris) 1948 (14): 23–28, six unnumbered figures.
Boiteau, P. & Mannoni, O. (1948b [juillet-septembre]). Les Kalanchoe (suite). [§ 3. Centrales (continued); Kalanchoe miniata Hils. & Bojer ex Tul.; Kalanchoe manginii Raym.-Hamet & H.Perrier; Kalanchoe jongmansii Raym.-Hamet & H.Perrier; Kalanchoe bergeri Raym.-Hamet]. Cactus (Paris) 1948 (15–16): 37–42, five unnumbered figures.
Bonnier, G.E.M. (1921 [January]) Flore complète illustrée en couleurs de France, Suisse et Belgique (comprenant la plupart des espèces d’Europe). Vol. IV. E. Orlhac, Paris; Delachaux et Niestlé, Neuchâtel; & J. Lebègue et Cie, Bruxelles, 134 pp., one Index page, 60 plates, [Plates 181–240]. [https://bibdigital.rjb.csic.es/en/records/item/15902-flore-complete-illustree-en-couleurs-de-france-suisse-et-belgique-t-4]
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Govaerts, R. (Comp.) (1996) World checklist of seed plants. Vol. 2. Part 1. Continental Publishing, Antwerp, Belgium, 492 pp.
Hamet, R. (1907) Monographie du genre Kalanchoe. Bulletin de l’Herbier Boissier, sér. 2, 7: 869–900. [https://www.biodiversitylibrary.org/item/105267#page/939/mode/1up]
Hamet, R. (1908) Monographie du genre Kalanchoe (Suite et fin.). Bulletin de l’Herbier Boissier, sér. 2, 8: 17–48. [https://www.biodiversitylibrary.org/item/104945#page/31/mode/1up]
Hamet, R. (1910 [d’Avril]) Kalanchoe aliciae sp. nova et K. beharensis Drake del Castillo. Bulletin de la Société Botanique de France [Tome Cinquante-Septième], Quatrième serie—Tome X) 57: 191–194. https://doi.org/10.1080/00378941.1910.10832196
Koorders, S.H. (1919 [Mai] [24 October 1918]) Beitrag zur Kenntniss der Flora von Java No. 18. Beschreibung von Kalanchoë Schumacheri vom Idjen-Plateau und Revision der Crassulaceae von Java. (Mit 2 lithographischen Tafeln.). Bulletin du Jardin Botanique de Buitenzorg sér. 3, 1 (3): 169–180, Figures 14 and 15. [https://www.biodiversitylibrary.org/item/27173#page/205/mode/1up (text); https://www.biodiversitylibrary.org/item/27173#page/243/mode/1up (figures)]
Mannoni, O. & Boiteau, P. (1947) Quatre Kalanchoe nouveaux de Madagascar. Notulae Systematicae [Herbier du Muséum de Paris. Phanérogramie. Paris] 13: 149–154. [https://www.biodiversitylibrary.org/item/7377#page/431/mode/1up]
Miller, P. (1754 [October]) The Gardeners Dictionary: containing the methods of cultivating and improving all sorts of trees, plants, and flowers, for the kitchen, fruit, and pleasure gardens, as also those which are used in medicine : with directions for the culture of vineyards, and making of wine in England : in which likewise are included the practical parts of husbandry [...] Vol. 2, 4<sup>th</sup> edn, corrected and enlarged. Printed for the author; and sold by John and James Rivington, London, pages unnumbered. https://doi.org/10.5962/bhl.title.79061
Perrier de la Bâthie, [J.M.]H.[A.] (1924 [1922–1923]) Les Crassulacées Malgaches. [Observations sur quelques espèces du genre Kalanchoe, pp. 21–24; Crassulacées Malgaches nouvelles, pp. 24–26; Stations et distribution géographique, pp. 26–28; Synopsis des Crassulacées Malgaches pp. 29–36.] Bulletin de l’Académie Malgache [Nouvelle série Tome VI] 6: 21–36. [https://www.biodiversitylibrary.org/item/50784#page/80/mode/1up]
Perrier de la Bâthie, [J.M.]H.[A.] (1928 [Février]) Observations nouvelles sur le genre Kalanchoe. Archives de Botanique II, Bulletin Mensuel. Caen 2 (2): 17–31.
Rouy, G. & Camus, E.-G. (1901 [Novembre]) Flore de France; ou, Description des plantes qui croissent spontanément en France, en Corse et en Alsace-Lorraine. Vol. 7. G. Rouy, Asnières (Seine) & E.-G. Camus, Paris, 406 pp. https://doi.org/10.5962/bhl.title.11986
Salisbury, R.A. (1805 [1 June 1805]) The Paradisus Londinensis: or coloured figures of plants cultivated in the vicinity of the metropolis 1. William Hooker, No. 6, Frith Street, city not stated, presumably London, 157 pp. & 117 plates. https://doi.org/10.5962/bhl.title.53520
Smith, G.F. (2020a) The kalanchoes (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1890–1972). Bradleya 38: 120–140. https://doi.org/10.25223/brad.n38.2020.a15
Smith, G.F. (2020b) Kalanchoe Adans. subg. Fernandesiae Gideon F.Sm. (Crassulaceae subfam. Kalanchooideae), a new subgenus from southern and south-tropical Africa. Haseltonia 27: 2–7. https://doi.org/10.2985/026.027.0101
Smith, G.F. & Figueiredo, E. (2018) The infrageneric classification and nomenclature of Kalanchoe Adans. (Crassulaceae), with special reference to the southern African species. Bradleya 36: 162–172. https://doi.org/10.25223/brad.n36.2018.a10
Smith, G.F., Klein, D.-P. & Shtein, R. (2021b) A revision of the Malagasy Kalanchoe subg. Kitchingia (Crassulaceae subfam. Kalanchooideae): history, taxonomy, and nomenclature. Phytotaxa 507 (1): 67–80. https://doi.org/10.11646/phytotaxa.507.1.3
Smith, G.F., Shtein, R. & Klein, D.-P. (2021a) Kalanchoe subg. Alatae (Crassulaceae subfam. Kalanchooideae), a new rank for K. sect. Alatae. Phytotaxa 509 (2): 255–258. https://doi.org/10.11646/phytotaxa.509.2.9
Thiers, B. (2020 [continuously updated]) Index herbariorum: a global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/ih/ (accessed November 2020)
Tulasne, L.-R. [‘Edmond’]. (1857) Fragmentum alterum (1). Floræ madagascariensis. II. Calanchoe [sic, Kalanchoe] pp. 149–150. Annales des Sciences Naturelles; Botanique série 4, 8: 44–163. [https://www.biodiversitylibrary.org/item/129817#page/153/mode/1up]
Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Oberreifenberg. [Regnum Vegetabile 159]. https://doi.org/10.12705/Code.2018
Baker, J.G. (1881 [21 February]) Notes on a collection of flowering plants made by L. Kitching, Esq., in Madagascar in 1879 (Plates VII. & VIII.). Journal of the Linnean Society, Botany 18: 264–281. https://doi.org/10.1111/j.1095-8339.1881.tb01257.x
Baker, J.G. (1882) Contributions to the flora of central Madagascar (to be continued). Journal of Botany, British and Foreign. London 20: 109–114. [https://www.biodiversitylibrary.org/item/35873#page/125/mode/1up]
Baker, J.G. (1887) Further contributions to the flora of Madagascar. Journal of the Linnean Society, Botany (London) 22: 441–537. https://doi.org/10.1111/j.1095-8339.1887.tb00472.x
Berger, A. (1930 [Mai]) Crassulaceae. Unterfam. II. Kalanchoideae [sic, Kalanchooideae]. 6. Kalanchoe Adanson. 7. Kitchingia Baker. 8. Bryophyllum Salisb. In: A. Engler & K. Prantl (Eds.) Die natürlichen Pflanzenfamilien. ed. 2, 18a. Verlag von Wilhelm Engelmann, Leipzig, pp. 402–412.
Boiteau, P. & Allorge-Boiteau, L. (1995). Kalanchoe (Crassulacées) de Madagascar. Systématique, écophysiologie et phytochimie. ICSN, CNRS, 91198 Gif-sur-Yvette, Éditions Karthala, Paris, 252 pp.
Boiteau, P. & Mannoni, O. (1948a [mars-juin]). Les Kalanchoe (suite). Section II. Bryophyllum. [§ 3. Centrales; Kalanchoe pseudocampanulata Mannoni & Boiteau; K. rolandi-bonapartei Raym.-Hamet & H.Perrier; K. pubescens Baker] (à suivre). Cactus (Paris) 1948 (14): 23–28, six unnumbered figures.
Boiteau, P. & Mannoni, O. (1948b [juillet-septembre]). Les Kalanchoe (suite). [§ 3. Centrales (continued); Kalanchoe miniata Hils. & Bojer ex Tul.; Kalanchoe manginii Raym.-Hamet & H.Perrier; Kalanchoe jongmansii Raym.-Hamet & H.Perrier; Kalanchoe bergeri Raym.-Hamet]. Cactus (Paris) 1948 (15–16): 37–42, five unnumbered figures.
Bonnier, G.E.M. (1921 [January]) Flore complète illustrée en couleurs de France, Suisse et Belgique (comprenant la plupart des espèces d’Europe). Vol. IV. E. Orlhac, Paris; Delachaux et Niestlé, Neuchâtel; & J. Lebègue et Cie, Bruxelles, 134 pp., one Index page, 60 plates, [Plates 181–240]. [https://bibdigital.rjb.csic.es/en/records/item/15902-flore-complete-illustree-en-couleurs-de-france-suisse-et-belgique-t-4]
Descoings, B. (2003) Kalanchoe. In: U. Eggli (Ed.) Illustrated handbook of succulent plants: Crassulaceae. Springer-Verlag, Berlin, pp. 143–181.
Govaerts, R. (Comp.) (1996) World checklist of seed plants. Vol. 2. Part 1. Continental Publishing, Antwerp, Belgium, 492 pp.
Hamet, R. (1907) Monographie du genre Kalanchoe. Bulletin de l’Herbier Boissier, sér. 2, 7: 869–900. [https://www.biodiversitylibrary.org/item/105267#page/939/mode/1up]
Hamet, R. (1908) Monographie du genre Kalanchoe (Suite et fin.). Bulletin de l’Herbier Boissier, sér. 2, 8: 17–48. [https://www.biodiversitylibrary.org/item/104945#page/31/mode/1up]
Hamet, R. (1910 [d’Avril]) Kalanchoe aliciae sp. nova et K. beharensis Drake del Castillo. Bulletin de la Société Botanique de France [Tome Cinquante-Septième], Quatrième serie—Tome X) 57: 191–194. https://doi.org/10.1080/00378941.1910.10832196
Koorders, S.H. (1919 [Mai] [24 October 1918]) Beitrag zur Kenntniss der Flora von Java No. 18. Beschreibung von Kalanchoë Schumacheri vom Idjen-Plateau und Revision der Crassulaceae von Java. (Mit 2 lithographischen Tafeln.). Bulletin du Jardin Botanique de Buitenzorg sér. 3, 1 (3): 169–180, Figures 14 and 15. [https://www.biodiversitylibrary.org/item/27173#page/205/mode/1up (text); https://www.biodiversitylibrary.org/item/27173#page/243/mode/1up (figures)]
Mannoni, O. & Boiteau, P. (1947) Quatre Kalanchoe nouveaux de Madagascar. Notulae Systematicae [Herbier du Muséum de Paris. Phanérogramie. Paris] 13: 149–154. [https://www.biodiversitylibrary.org/item/7377#page/431/mode/1up]
Miller, P. (1754 [October]) The Gardeners Dictionary: containing the methods of cultivating and improving all sorts of trees, plants, and flowers, for the kitchen, fruit, and pleasure gardens, as also those which are used in medicine : with directions for the culture of vineyards, and making of wine in England : in which likewise are included the practical parts of husbandry [...] Vol. 2, 4<sup>th</sup> edn, corrected and enlarged. Printed for the author; and sold by John and James Rivington, London, pages unnumbered. https://doi.org/10.5962/bhl.title.79061
Perrier de la Bâthie, [J.M.]H.[A.] (1924 [1922–1923]) Les Crassulacées Malgaches. [Observations sur quelques espèces du genre Kalanchoe, pp. 21–24; Crassulacées Malgaches nouvelles, pp. 24–26; Stations et distribution géographique, pp. 26–28; Synopsis des Crassulacées Malgaches pp. 29–36.] Bulletin de l’Académie Malgache [Nouvelle série Tome VI] 6: 21–36. [https://www.biodiversitylibrary.org/item/50784#page/80/mode/1up]
Perrier de la Bâthie, [J.M.]H.[A.] (1928 [Février]) Observations nouvelles sur le genre Kalanchoe. Archives de Botanique II, Bulletin Mensuel. Caen 2 (2): 17–31.
Rouy, G. & Camus, E.-G. (1901 [Novembre]) Flore de France; ou, Description des plantes qui croissent spontanément en France, en Corse et en Alsace-Lorraine. Vol. 7. G. Rouy, Asnières (Seine) & E.-G. Camus, Paris, 406 pp. https://doi.org/10.5962/bhl.title.11986
Salisbury, R.A. (1805 [1 June 1805]) The Paradisus Londinensis: or coloured figures of plants cultivated in the vicinity of the metropolis 1. William Hooker, No. 6, Frith Street, city not stated, presumably London, 157 pp. & 117 plates. https://doi.org/10.5962/bhl.title.53520
Smith, G.F. (2020a) The kalanchoes (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1890–1972). Bradleya 38: 120–140. https://doi.org/10.25223/brad.n38.2020.a15
Smith, G.F. (2020b) Kalanchoe Adans. subg. Fernandesiae Gideon F.Sm. (Crassulaceae subfam. Kalanchooideae), a new subgenus from southern and south-tropical Africa. Haseltonia 27: 2–7. https://doi.org/10.2985/026.027.0101
Smith, G.F. & Figueiredo, E. (2018) The infrageneric classification and nomenclature of Kalanchoe Adans. (Crassulaceae), with special reference to the southern African species. Bradleya 36: 162–172. https://doi.org/10.25223/brad.n36.2018.a10
Smith, G.F., Klein, D.-P. & Shtein, R. (2021b) A revision of the Malagasy Kalanchoe subg. Kitchingia (Crassulaceae subfam. Kalanchooideae): history, taxonomy, and nomenclature. Phytotaxa 507 (1): 67–80. https://doi.org/10.11646/phytotaxa.507.1.3
Smith, G.F., Shtein, R. & Klein, D.-P. (2021a) Kalanchoe subg. Alatae (Crassulaceae subfam. Kalanchooideae), a new rank for K. sect. Alatae. Phytotaxa 509 (2): 255–258. https://doi.org/10.11646/phytotaxa.509.2.9
Thiers, B. (2020 [continuously updated]) Index herbariorum: a global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/ih/ (accessed November 2020)
Tulasne, L.-R. [‘Edmond’]. (1857) Fragmentum alterum (1). Floræ madagascariensis. II. Calanchoe [sic, Kalanchoe] pp. 149–150. Annales des Sciences Naturelles; Botanique série 4, 8: 44–163. [https://www.biodiversitylibrary.org/item/129817#page/153/mode/1up]
Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Oberreifenberg. [Regnum Vegetabile 159]. https://doi.org/10.12705/Code.2018