Phephu, N., E. T. Witkowski, J. V. Rooy, and M. Sim-Sim. “An Updated Checklist of the Liverworts and Hornworts of Southern Africa”. Phytotaxa, vol. 725, no. 3, Oct. 2025, pp. 231-70, doi:10.11646/phytotaxa.725.3.2.