Heinrichs, J.; Dong, S.; Schäfer-Verwimp, A.; Peralta, D. F.; Feldberg, K.; Schmidt, A. R.; Schneider, H. Towards a Monophyletic Classification of Lejeuneaceae II: Subtribes Pycnolejeuneinae and Xylolejeuneinae Subtr. nov., Transfer of Otolejeunea To Lepidolejeuninae, and Generic Refinements. Phytotaxa 2014, 163, 61–76.